Taxonomy and Fieldwork

James Deacon asked Bruce Bayer, “What is this thing at Brandrivier?”

James – you have no idea how significant this picture is. It is an emelyae variant and I really need to know a lot more about precise whereabouts. Multifolia comes from a few km to the east and there was a very very rossouwii-like multifiolia on Brandrivier. It is pictured in Revisited. But it is not there anymore. What is there are plants like your picture indicate. Did you perhaps observe Tulista opalina?

Jakub Jilemicky added “This plant was described by Gerhard Marx as H. obserata. It occurs mainly at Brandrivier, but as well at Springfontein. It flowers with all the magnifica type plants, not with emelyae group.”

May 19 – 7888 Brandrivier. H. emelyae ‘obserrata’. It raises an interesting issue. Names may have to change every generation as the plants may change? It used to be ‘multifolia‘. Actually my species definition includes the matter of space and time. In which case the name ‘obserrata‘ is taxonomic malfeasance.

May 19 – 7846 Between Springfontein and Brandrivier. Flowering August. Surely the type loc for “H. obserrata” that I see as H. emelyae. But this is just a distraction. When you still have commentators making comparisons to the magnifica types and flowering times, you realize that the absurdity of Haworthia classification is going to continue for the next 80 years as well.

As a point of interest. There seem to be many different ideas of what species are. From the typological concept of things that look near identical, to a list of the species one thinks makes up a genus. In the latter case the genus is considered to be what is made up from the species one imagines. My own species concept is very well defined and my map (this spider web of colours) indicates the way factors of space and time are involved in only one small group of Haworthia.

Still paradoxa Vermaaklikheid but the first few were June and this is September.

Let me try this as no 3…What we have is a post of three pictures to show three “species” paradoxa, bobii and joleeniae and to ridicule Bayer for suggesting they are the same. Firstly those pictures are NOT the species. They are specimens taken to illustrate the species but by what is recognised in botanical taxonomy as the “typological concept”, i.e. a species consists of individuals that all look the same. I wish I could copy here what I wrote in the New Haworthia Handbook (1976) where I use the name H. magnifica var. paradoxa and discuss affinity with H. turgida, H. emelyae, H. retusa and H. mirabilis. I ended the discussion there with this… Consideration of the variability of this species and distribution of variants, is very helpful towards understanding variability in the genus as a whole. What I conclude is that the species are systems with any number of populations and individuals that vary enormously. The typological concept creates mayhem in the minds of critics, growers and collectors alike. So let us start with the paradoxa, but first dismiss H. magnifica as a species. That name is a total myth. The myth ghosts around probably a single plant I collected with J. Dekenah at the type locality. It proved very amenable in cultivation and plants ended up at Sheilam nursery to go world-wide. It is actually difficult to now find a clone quite the same at the original locality where there are several discrete “populations” in quite a small area – and countless variants. It is senseless to say that H. magnifica can be a species if in every aspect of such a decision it is topologically based, as is the unfortunate case. The name in SANBI is H. mirabilis. Paradoxa is a synonym. SANBI are not my friends or my clients. Some very bright and competent people are employed there. The element paradoxa is thus in my view a variant in the species H. mirabilis and illustrated here by three populations viz. at Vermaaklikheid, Osplaas and Koenserus.

Just get something clear about type specimens and the system. A specimen is only anchor for the name. It does not prove anything about where it comes from. It is just representative. A second point is in that the nomenclatural rules uphold chronological discovery (better still – description) over reality. This mad rule of priority totally confounds the prime aim of a classification that follows and explains the genetic history (phylogeny). If in fact evolutionary theory is even valid. Generally the rules provide fertile ground for confusion, argument and publication credits that do little to promote their purpose.

Two Vermaaklikheid picture sets have been posted. This is another. A cooler west facing outlook but also on those limestone rocks. I must confess to a bit of apprehension that some may claim this is a different species when my opinion will be a great deal more conservative in what lies ahead.

What is the problem? I suggest we have an antiquated, outdated, dysfunctional classification system so that our knowledge, philosophy and methodology does not allow explanation of groups like Haworthia. So I will try and demonstrate it like this…Gerhard posted pictures of paradoxa, bobii and joleeniae. One of each! So let us just see how this can be interpreted. Here are several pictures of ‘paradoxa‘ from Vermaaklikheid east of the village. I first saw it at a site north of the village.

There is so much still to explore. Here is a map just to indicate the scale that has to be exercised. Initially I was happy to stop every 50 miles to see what might be there. Now it is every 25meters! Just note an interesting phenomenon here. 3 species and as many populations as there are recorded habitats. M = mirabilis, R retusa and F floribunda. Yes we covered most of the ground between too.

Mukesh Vaid asked – What does these four digit numbers signify

Bruce Bayer – Just my collecting numbers. This map is only intended to show intensity of search to demonstrate anything. I did have a collecting permit strictly adhered to. Often, as I do now, I do my searching with camera!

Every collector should have an accession book to record what you record and find where and when. I idiotically did not get this right to start with. Even as an entomologist I did not keep proper numbers. When I started at Karoo Garden I used the garden accession system KG with number/year. No good. In the end it is the quality of accession record that indicates the credibility of the work! I did not have the luxury of digital camera or GPS that is essential nowadays. At one time it was enough to just say “collected between the Cape and the diamond fields! With plant theft and cadaverdog-like shadows, it is a bit stressful to even produce the maps that I show! Sometimes I have mentioned something and the next moment a new species is described directly connected??

Mukesh Vaid – Where is H. turgida in this?”

Bruce Bayer – This not actually about turgida but just to show level of search necessary. But let me take the opportunity to try and get this message/hypothesis across. There is a single gene pool (a system) that produces mirabilis, retusa, turgida, pygmaea, mutica and emelyae. It can be simplified if it recognised that retusa is really a offset of turgida in among 5 main lines. So let us try and get something else straight. Science (i.e. scientism actually) is a system of reductionism that supposes that everything can be explained by breaking things down to their smallest parts. So we mess up in Haworthia by thinking the smallest parts are species when in fact it is a system of many things. Does that help?

 

This is where I think Mukesh rightly has a problem. That bottom pink line should not be there. That is why I refer to them as intervals. These bureaucratic scientists lay down the law that this should be seen for what it is and call it an interval – because it is a break in distribution continuity e.g.. The Knysna interval is a serious break between SE coast flora and S Cape flora. That coast line area is not easily explored and the geology is limestone with a different vegetation. There may well be Haworthias there. There is a record of a mirabiloid(?) on the sea cliffs at St Sebastian west of the Breede River mouth.

There is a really weird intimation that bureaucrats have determined what species names apply on Haworthia! It is actually a very democratic process based on 4 tenets of science. Universal truth, communality – no secrets, no private gain, and organised scepticism. If you have the data and you have the facts, you are free to organise and present your case. I have no privileged access to anything and certainly no sycophantic following with commercial interests in mind. I often wonder that not a hair sticks up from the trenches of my defense when so much garbage is thrown on attack

On this map…. There is actually no known connection across from L = longebracteata (also a mirabiloid area), to P = paradoxa, to B = bobii. Historically the shore line has changed and there may have been. Variegata seems to have jumped the gaps. So now we have populations a = Buffelsfontein, b = Sandfontein, c = Sandhoogte and d = Infanta. A and b are in the De Hoop Reserve and it is quite difficult to access so I have not been there with a digital camera unfortunately. Buffelsfontein I owe to Chris Burgers, Sandfontein to Adam Harrower and Sandhoogte to Jakub Jelimicky. Infanta I owe to mesemb research and what “bob” had to do with it is anybody’s guess. Certainly I did not remotely think it was or is a new species, as excited as I was to first see it. The overall complexity of the spider web does not do justice for the problems of variability among the floribundoids, the variegatoids and the mirabiloids. Turgida seems to hold its own. Holistically I can conceive of only the one solution that excludes adding another few species to an impossibly and unrealistic list. It would be nice if the culprits owed up to the chain of events that led to “bobii” and a few other names. Including the authorization for collecting that is so uncomfortable and dehumanizing. I make no apology for suggesting the association of bobii and paradoxa , even beyond a greater issue of the mirabiloids. I do not doubt that it is a difficult “ask”. The two pictures are of ‘a’ and do not do justice to the plants in the field at all.

Arthur Dixon posted these .. Bruce Bayer replied “All ‘paradoxa’.”

These are the plants at ‘b’ where Adam Harrower recorded them. I named the place as Sandfontein or “Sandhoogte N”. Makes me long to go and drool over them again!

These are my pictures of ‘c’ in the ‘bobii’/paradoxa/joleeniae/mirabilis milieu. Funnily enough I really do sympathise with the doubters and unbelievers and am very apprehensive about further showing you what I think is the same. (remind me if I forget!). I have not mentioned “muticoid” in this lot where I say turgida is real. I suggest that mutica does come in to the picture and I will touch on that when I get to ‘joleeniae’.

Arthur posts…”It seems to me that endless discussions and reasoned arguments supported by habitat shots whilst life-affirming and endlessly enthralling are not advancing the discussion in any way as (returns to main point) WITHOUT that definition of ‘species’ the cycle continues to revolve (Personal point)”. Yes yes yes. It has been ongoing for the last 80 years. Like I said – I get credit for my great contribution to Haworthia while effectively been told I know nothing. I am happy to accept that, if some intelligence is put in place of what I have written over the years. Contrary to general belief, I do not like offending or hurting people. Therefore I am very limited in what I can actually say and tell. What we need is a dimensional shift in how we think, how we reason and in what we are really wanting to achieve. We have a societal crisis and we see it not!

Tony Brook wrote “My apologies Bruce I am not a Botanist but surely the definition and description of Bobbi should precluded the 2 plants which are relatively hairless. Otherwise many other Haworthia would be included in the “species”.”

Bruce Bayer replied “No Tony. this is essentially the problem. I did not consider bobii to be a species. H. mirabilissii the species and the best way of defining species at the moment is by DNA sequencing but rather also by what my definition of species requires. Viz a holistic view of its relationships, similarities whatever, with possible and probable relatives.”

Here is another ‘joleneae‘ population that demonstrates how observations can be slanted to re-enforce propaganda. Someone also sent me an article on Haworthiopsis that is a piece of real modern day journalism. Unable to tell left from right or right from wrong, the writer sits on the fence like a Fiscal Shrike impaling worms on the barbs, i.e. hoping to have his believers (us ordinary worms) accept him as an all-seeing eye of truth. This has plagued me all my writing life! No joleneae is not definitely and distinctly, always glabrous. Some of these pictures even show the mutica shiny speckled appearance those plants can have. Differences between populations are in my observation as much due to the skeletal soil differences (geology) as they are to spatial isolation. I think. Some amazing individuals here.

So let us move on because there are still surprises and unpalatable truths. While we have seen mirabilis ‘bobii‘ from the sandstones of the Potberg itself, we need to see what mirabilis does on the geological confoundment of the flats north of the Potberg. First there is Melkhoutfontein and I assume that this is included in the concept of “joleneae” (that I usually misspell). It really should be H. mirabilis ‘joleneae‘ and just forget formal process. It is not possible to regulate and manage the welter of names. An attempt was made to register Aloe cultivars, but if this is practical I doubt. It is senseless to argue there is no continuity from ‘bobii‘ to ‘joleneae‘. This just upholds a nonsensical humongous heap of formal binomials to argue about and ignores the many more things that attract either no aesthetic attention or commercial interest. Joleneae is on the terrace cobbly sandstone as a simply descriptor.

It occurs to me. Joleneae is claimed to be consistently glabrous. This comment disregards the electron microscopy “work” done by Dr. David Cutler of Kew that I reported on in Updates. It is obvious from these pictures that the plants are incipiently hairy or spinose whatever is correct. we simply cannot properly assess the real nature of individual characters like this. Just for recall – there was an amazing paper written on the cladistics of the Aloids published in Taxon (about 30 years ago). Whoever peer reviewed that paper prior to publication deserves serious censure. Barely a character state is correctly stated.

What it needs to is for the thought leaders, like editors, society committee members, media notaries etc. to get their act together and seriously consider what their role is in the mess of argument and infighting. I have suggested now a solution where at least in Haworthia we can (if we so choose) admit the spider web and set-out how to communicate about it. Perhaps we need to identify the mafia who have vested interest or are just too intellectually lazy or impractical to even want agreement.

Is this and that getting too much for you? Still ‘joleneae‘. It is a useful name, but not actually the issue. At this place Tulista minima (I am gobsmacked that some over zealous mind has decided it is actually T. minor – I suppose this name change in intended to avoid confusion!!! ) as does H. variegata ‘hemicrypta’. So I include hybrid pictures. The two species do not directly share habitat. Stoffelsriver.

Here is a map of the north Potberg area to highlight the significance of geology and its impact on floristics/vegetation/plant species. It is a complex situation with deposited materials overlying basic formations and erosional effects. The white material to the higher left are eroded and decayed shales that result in banks of white clay much like elsewhere towards Riversdale and Heidelberg. Is it surprising that the retusa/mirabiloids adjust and adapt accordingly. Why do these plants differ so much from joleneae/bobii/paradoxa and switch to resemble atrofusca and floribunda?

Snakes and ladders. Background noise! I have 206 folders for populations of which most are this unphotogenic ilk. Their lack of appeal and hidden nature is why H. heidelbergensis was spawned. The more conspicuous and bigger forms were H. maraisii and H. magnifica. The only attention I know they ever got from collectors was the complaint that they never could own or get a proper idea of what H. mirabilis looked like. Shades of the great doyen of Botanical Latin and plant taxonomy W.T. Stearn who destroyed the foundation of Haworthia taxonomy with his typification of the name Aloe atrovirens, spinis herbaceus numerosus ornata and choosing the herbaceus part for the epithet. H. atrovirens should be H. mirabilis! I do not think the name mirabilis was even used in von Poellnitz and Smith’s time? So I can see that my chances of putting Haworthia taxonomy to rest are zero. No herbarium can ever hope to hold a proper record of variation within a single population. Whoever wants to quarrel with me needs to now that their classification includes all the stuff I have tried to contend with! My pictures of this background noise are disappointing and I have none for stuff seen before ca. 2005 or so. Emoji of laughter or tears? And who wants to know what a species is anyway?

Nothospecies or notaspecies? That is the big question. Why bother to ask difficult questions, just call this H. mirabilisXmutica H. hammeri instead. It consists of one population occupying an area less than 500sqm among surrounding mirabilis and mutica populations. Will it ever be a system? (Nothospecies is coined for the offspring of two other species that in the absence of a definition become a new species. It really applies to readily seed propagating and readily dispersing plants – agriculture and horticulture!)

Here is my 2009 baby I think written in 2009 before DNA sequencing proved to us that there were 3 genera where we saw only 1. I said so when? First handbook 1976? The thing that really got me is that the results of the sequencing were actually fudged (again). The discussion and conclusions were written long before the introduction and methods, to ensure that there was a match up of results to hypothesis being tested.

Haworthia the problem child of taxonomy

Yes my favorite critic had something to say about this fishtail bud as well. Or at least the fact that I considered it at all. It is actually a feature of virtually all of the retusoid mirabiloids varying from zero in the east to nearly dramatic in the west. So herbacea and reticulata have the most exaggerated version. In these two systems the flower is nearly “regular” (as in pentagonal) hence reticulata ‘subregularis’. H. herbacea is characterised by the beige colouring while reticulata flower is pinkish. The Wolfkloof Villiersdorp herbacea has an enormous (close to 3cm long) very pink flower to fudge the issue. H. herbacea ‘paynei’ near McGregor has a bicoloured flower, hence ‘luteorosea’.

6509 H. mirabilis meiringii, W Bonnievale – Flowers are problematic because the flowers are not static things. They change by the day. In this case I pictured most open flowers on every single stem from 1-5 open.

6089 NW Potberg – some of the background noise to mirabilis. Many many populations of these small black things (“klein swartetjies” according to Michael Malherbe founder of Sheilam, in imitation of the Little Brown Jobs of bird-watching)). Not as enchanting as the larger and less common variants, they are not covered by popular classification.

H. mirabilis ‘notabilis’. This is a place in the mountains midway between Robertson and Worcester. Both arachnoidea and reticulata occur here as well in different habitats. There are shale, dolomite and sandstone formations. The countryside is formidable. How does one describe the plants here and link them adequately to a host of populations running all the way east to Ashton, and to the oddities of the NE Worcester area. Not only that. There are continuities southwards as well.

Lets try something. First the chaos formula imaging. Like a DNA seq. phylogram it is two dimensional and cannot adequately represent a spacetime issue (that species are whether evolutionary or independent creation of ‘species’. Secondly imagine a continuation of the fractal image with 6 species in a particular time frame continuing on to eventually end up as 3 species in a third frame? The frames are geographic space. The brackets demonstrate how species may overlap in distribution and/or characters. The chaos formula demonstrates overlap of characters in time or space but not in both as is needed. That is called linearity as opposed to actual reticulate reality? I am no intellectual wizard but am quite sure that plant taxonomy has got things WRONG.

Guan Tan-Amo Lim wrote “Haworthias are a little bit like domesticated dogs… they come in all shapes and sizes and can breed with each other readily. Yet dogs are all of one species. Could the many Haworthia species similarly be reduced to if not one species then to only a few? With many “breeds” corresponding to the natural populations in habitat?”

Bruce Bayer replied “Guan Yes. but we shift the problem from one level to another. I have a spider web of five species (turgida/retusa, emelyae, pygmaea, mutica, emelyae that could be one. But there is another spider web in the upper left of herbacea, reticulata, maculata and pubescens. So where do we stop? Things like wittebergensis, pulchella and marxii seem totally separate but there is chaos among all the other true Haworthia. Funny I was just browsing an article on a math issue that claims that there are things that are true but they cannot be proved to be true? Life is a conundrum.”

A fairly random lot? Actually no. This is a place a few km from ‘hammeri‘. Their is a rounded hill with small nondescript LBJs (little black jobs) most places, but at one spot quite large plants like this. Atrofuscoid? The floribunda-like leaf tip is common in the small mirabiloids. I should add… a note on fieldwork. Did I do field work? Walking around with nothing to measure and itemise but an eyeball assessment and some pictures.

6638 from NNE Bredasdorp. Mirabilis. This is a place where turgida also occurs, and with rossouwii and mutica within a few km. Also home to the ‘heidelbergensis minor’ that I suggest has rossouwii genetic material. One of these clones is very reminiscent of what I observed and considered a true hybrid between mutica and mirabilis where populations were in very close association. Does anyone really have problems with names used in an informal way like this rather than behind a facade of deep science and a degree of fraud?

From 6638 pictures. This weirdo strikes me as reminiscent of more obvious hybrids retusa/mirabilis or mutica/mirabilis where they are in close enough proximity to support a hybrid presumption. Turgida is an estimated 400m away from here.

Soumen Aditya comments “I didn’t see a features of natural hybrid … Most probably the polymorphic evolution…” Here is what IS most probably hybrid. Where mirabilis and mutica are in two populations meters apart. SO I think what would be features of a natural hybrid may just be a subjective opinion? I have mentioned before how problematic the issue of hybrids is and have often been asked – How do you tell it is a hybrid? Credibility in classification barely exists at the best of times.”

What do I ask? I once was in a meeting with a group of top managers discussing a topic they had been dealing with for over 20 years viz. rangeland production. I was suggesting a method of assessing potential. The group seemed so lukewarm to my suggestion that I felt the need to establish if they really did know what I was talking about. So I sent around a problem and asked if it could be solved using the available knowledge they had been debating for over the previous 20plus years. Deadly quiet. No wonder they were so indifferent. Now I feel exactly the same about Haworthia classification. People have been arguing and debating this since Herre’s time (1930). Here in SA nothing has changed. Perhaps I also do not know what I am talking about! The spider web for the retusoid/mirabiloid plants is a small part of the whole, and fairly straightforward compared to the bigger picture. But my suggestion for a systems approach is met with the same deadly quiet, especially from where it matters. What do I do next?

These are a few pictures of 30 from a population in a vast expanse of a no go area for any but the most intrepid. What is it? It is from an aloid sparse space and while I do have pictures for another 5 populations in the area they are not more informative. From my limited (is it that bad?) experience of mountains and the way succulent exploration has been done, I shudder to think how the Haworthia community, led as it is, is going to be able to digest what is still to be learned.

I mentioned that vast no-go area (6694). Here is one of the other 5 populations I know that happens to be only about 500m away (7994).

7994 Yes the flowers are significantly different. 7994 I attributed to maculata and the fishtail bud indicates that it is an extension of the retusa/mirabiloid web. The rounded bud tips of 6694 suggest arachnoidea or nortieri, as geographic candidates and the most probable system affinities.

June 4, 2021 – Something useful I can do is amplify H. mirabilis meiringii. I described this to draw attention to very herbacea like forms of mirabilis. This is in the Bonnievale area where the mirabilis variants seem like a system within a system, within a system. Westwards they are less spinescent and similarly eastwards. 6509. The name was not to honour someone so much as to record a time when plants were perhaps first harvested and sold from the field.

Silly me. I say these are the same (species)

H. mirabilis meiringii 7269

Mirabilis meiringii 7327

meiringii 7882

H. mirabilis Klipfontein

 ♦

Zuurberg cooperoid

I have seen about 35 Zuurberg cooperoid populations in the vicinity of the Kaboegapoort, N Kirkwood. None of these are quite the same and reflect an interaction of at least 4 megaspecies (in this case decipiens, cymbiformis, cooperi, bolusii (blackbeardiana) and aristata. Please note – it is taking a conservative middle ground opinion to say they are discrete!!! This is MBB6940 and the variation in just one of the Kaboega populations. Do not let it be lost on you that it was (and more than probably is still the case) that a species was too often described from a single specimen. Think of Rafinesque. There is absolutely no way these things can be regarded as distinct species in their own right. I have seen too many of these local variants linked not to just geographical separation, but also to local habitat difference and usually geologically associated.

7870 and 7586 It will have some dinosaurs climbing the wall when I say the probability of these both being H. cooperi is 98%. Unfortunately it is TRUE. the bobii, paradoxa, joleneae, hammeri, groenewaldii issues are minor compared with this. Closed minds will struggle and someday taxonomic botany itself is going to have to face not having a proper species concept and definition. The Haworthia community would do well to properly acknowledge the fact that it has a problem.

Lawrence wrote … And the problem is these Zuurberg cooperoid need names?
Bruce relied … They sure do. The science of botany is not going to do it and we the people have to fix it. Trouble is we reduce the issue to personality disorders around the issue of Latin binomials instead of applying our energy to thinking communally of how to find a solution. ♦

Maculata

A whole bunch of pictures of H. maculata. I would have thought my Handbooks might have discouraged the “new species” guys who do not seem to recognise what the nature of the game is. There were about 450 names being bandied about that I reduced to about 140. It just seemed to spur these guys on to inventing several hundred more to recreate an unmanageable situation? Perhaps the science of botany itself lost the plot when it was decided that the practicality of genera was actually built around the need for management rather than a dubious science of evolution.

Clump

Steven Molteno – Could almost imagine these on a steep shale riverbank in the Overberg.

Bruce Bayer – Aren’t these clump formers usually associated with riverine or stream defined rock slopes and faces? While maculata has the options to be a solitary plant or a clump former. Herbacea can be a clump former in the absence of a watercourse while at Wolfkloof in a gorger situation it is fairly non-clumping. I come to insist that science has truly led us away from understanding anything about a conscious creation. Mechanistic answers do not hold truth and we are not open enough to be aware of it?

Steven Molteno – True. Same with Tulista I guess, comparing minima growing large & solitary on hills (“retusa-minimas”!) vs. minima growing in clumps along river-banks (“turgida-minimas”!) I’ve become interested in which of the Haworthia northerners might be (relatively) closest to the retusoid grouping of the Overberg. Reticulata has many analogies with the river-born retusoids (turgida), not least in terms of habitat/substrate etc. The translucent grouping of mucronata/cooperi etc. sometimes seems to show similarities to them too maybe? If it’s not my imagination. I’ve even seen photos of some cooperi/cymbiformis in the eastern cape that I could easily mistake for chubby turgida of the Overberg. But going in a different direction. I wonder about the nortieri/marumiana groupings and whether they link to the retusoids too, via maculata perhaps.

Bruce Bayer – Of course – there is no doubt in my mind that something separated off reticulata/herbacea/maculata from the retusa/mirabilis lot. I have also considered the possibility that maculata is the link of that combined system to nortieri. There is a mucronata on the Anysberg pass slopes that is inseparable from cymbiformis except flower wise. So it gets extremely convoluted because I can see cymbiformis and cooperi as a single system – inseparable from mucronata – inseparable from arachnoidea. Marumiana seems to have its ties to monticoloa – to then angustifolia, while flowerwise angustifolia and variegata are inseparable. There are just tons of missing pieces (and not so missing pieces). We just get sidetracked by truly absurd opinions about things like ‘groenewaldii’ – as a plum-picking opportunity by those with heads in the sand!!!

Steven Molteno – The multitude of links in the Haworthia lattice can leave one quite dizzy. I’ve sometimes found it helpful to (over)emphasise to myself the different types of linkage. Mainly, the distinction between confluences (ie. links through hybridisation) and divergences (ie. links through integradation). Put simply – coming together vs. moving apart. It’s one of the few ways I can get any sort of picture in my head, as we’re probably dealing with a multi-dimensional lattice of clines that’s nearly impossible to untangle. I’d be interested to hear your thoughts on the strength & nature of the link between: nortieri (+maculata etc.), and marumiana (+archeri etc.)

Bruce Bayer – Both those relationships are really speculative and they revolve around the oddities of the Hex Valley and eastwards as far as SE Laingsburg. There is just not enough information from which to draw any better conclusion. As you know from Astroloba, that area holds mystery and H. marxii contributes, but so does pulchella and wittebergensis. Dimorpha??? There are at least 5 populations assigned to marumiana in the western Karoo and then there is the problem of Reddii in the east. None of this is adequately supported in the way that can be argued for the retusoid synthesis. Nearly every botanist exploring these higher points in the Karoo fail to report a marumiana-like presence.

Steven Molteno – That area does indeed hold a lot of Easter eggs. To mix the metaphor, the Wagendrift formation especially seems to be a museum of weird relics. H. marxii is almost entirely confined to some narrow strips of Wagendrift rock; H. wittebergensis is mostly (not entirely) also on this formation, and that new plant recently found (H. grenieri I think it’s called) is too of course. I’ve no idea why that’s the case though.

So would you agree that, in terms of relationships between groupings:

  1. H. maculata is probably closer to nortieri (relatively!) than it is to marumiana?
  2. That archeri is more closely linked to marumiana than to nortieri? I do seem to recall reading in one of the Updates that there are also (weaker?) links between archeri and both nortieri and pulchella (or have I imagined that?)
  3. That there is potentially also a connection between the nortieri group and the marumiana group, but that this is highly speculative as you say?

Okay I think I understand what you mean:

  1. nortieri cannot be connected both to maculata and to marumiana (as you say, diametrically opposed), so you believe it must be either one connection or the other, but not both. Based on your earlier comment then, I assume that you posit the connection only to be: maculata to nortieri (if a connection there is)
  2. Understood. Though I cannot help seeing similarities between pulchella and herbacea (& rossouwii)
  3. Well, your answer to 2. serves as an answer to 3. as well.
    Afterthought … I do know how unreliable flowering time can be, as a diagnostic criterion for tweezing apart Haworthia taxa, so that’s fully acknowledged. But it’s still a bit interesting that some of these northerners are +-Spring flowering, e.g. nortieri (Sept-Oct?),maculata (Sept-Oct-Nov?), notabilis (Sept for some populations?) and that marumiana is apparently late-Summer flowering (Jan-Feb), just like the variegata-floribunda grouping.

Bruce Bayer – No, maculata to nortieri and nortieri to marumiana are diametricaly opposed.
Yes to first part. The archeri nortieri link would be via Hex Pass and very weak. Pulchella simply stands alone. 3.What do you mean nortieri group? when already norieri to marumiana is via archeri? I think we could start again from scratch. We are trying to establish what the systems are – if you now have decided what you think a species might be. Nortieri appears to me to be an independent system. There is a possibility of some connection to arachnoidea although they do occur in very close proximity in specific instances. The Hex valley plants just means I can’t exclude other possibilities. I see no connection between herbacea and pulchella and the flowers of the former place it firmly with the retusoids. There is no doubt rossouwii comes into the retusoid picture but indirectly. I would guard against trying to hard to establish connections like this other than as a driver to find evidence to support the process. I suggest you clear your own mind about “species” and recognise too that a mechanistic answer may not be available. ♦

A random note about Haworthia chlorocantha

Haworthia chlorocantha is a name I use to cover a whole hoard of populations of plants that occur in the Southern Cape south of the Langeberg mountain range between George and Albertinia. Of course, as is the despairing fact in Haworthia, the plants in the different populations or even within populations, often do not look the same at all. It is only from a distance and from an overview of all the available (note ”available” – to me?) evidence, that I concluded that there are two sets of plants in the area. One is the group that I refer to as the Retusoids. It comprises H. retusa (=H. turgida), H. mirabilis, H. pygmaea, H. mutica and H. emelyae), although technically only H. pygmaea and “H. turgida” are in the designated area. The other group is the floribundoids that are H. floribunda, H. chlorocantha, H. parksiana, and H. variegata. Quite rightly and understandably, opinions about this as regards species names, differ humungously. I can only explain what my experience and information suggest. I lump the names like this because there are undeniable problems about regarding them as made up of different as fully and clearly different systems we like to name as species. Furthermore I must add that the problem is compounded when one is faced with the reality that for example H. floribunda, interacts in the field with the retusoids as well as with its fellow floribundoids.

Here I would like to just show and discuss one population of Haworthia chlorocantha that I recently came across east of Albertinia in a twilight zone where its relationship to H. floribunda is questionable. I also grew plants from seed and I will show what they look like after two years. This is particularly interesting because Essie Esterhuizen has done some remarkable Haworthia exploration over the last 40 years and commented to me upon his inability also to not find any real dramatic and clear differences within the same two groups I mention. His is a totally independent observational route to my own and he might well express his thoughts quite differently and understandably even choose to do so. Unfortunately I just do not have a picture record of all the Haworthia chlorocantha populations (and of course the others too) that I have personally seen. I would so like enthusiasts to have that available to them. So this short essay is restricted to just this one single locality and disregards others.

I have not even allocated an accession number to this population north east of Albertinea along the Valsch river. There is another even more complex population still further east that Etwin Aslander found and on which I commentated on in my Haworthia Updates. My first five figures are of adult plants in a small localized population occupying about 15sq m in Renosterveld. As you can see the plant are quite variable and there is undoubtedly I floribunda similarity in the leaves. Thinking that maybe flower characters may hold some significance is just a joke as I explained in Vol 8 of Updates. This applies nearly equally to flowering time that is March for this population.

Fig. 6 is of two pots of seedlings. Repotted at one year, they separated into two size groups. This is in itself an indication of internal population difference and my personal experience of growing these Haworthias from seed is that there is a natural inclination to first pick out the more robust seedlings and work down to the small weakest, sometimes even discarding the last!  Even in this case I have discarded seedlings. What did I lose?  So at two years we have a pot with 4 plants in the size range of diameter spread up to 4.5cm (figs. 7 and 8) and another pot with the spread up to only 3.5cm (fig. 9 showing a plant with a 2cm leafspread. There are differences in colour and leaf spination. With active growth in shadow, the leaves are recurved. Some of the leaves show the characteristic twisted and rounded tips of H. floribunda. The smallest plant is indicative of the H. parksiana-like size that suggests the same thoughts of species similarity that come to mind when exploring other populations.

published in Haworthiad v36.1 pp 14-16, February 2022 ♦

Tulista

A series of Facebook postings from April & May 2022 covering Tulista kingiana, T. marginata, T. minima (minor), T. pumila.

Warwick sent me this great picture of Tulista pumila taken between Worcester and Robertson. It would be interesting to know how long-lived individual plants are because this species can form an expanding ring of offsets much like Aloe claviflora does.

Nuisance – prempting Warwick’s post was a disaster. This is what Shucked should be about. A celebration of nature and what she has for us. We are in a mess because we have no respect for nature or for its many life forms. If it moves we shoot and kill. If it grows we chop it out or down and make charcoal, choke medical properties out of it to cure the many ailments we get due to maltreatment of other life forms? Do we collect and propagate Haworthia for the sheer joy of nature or because we get some sort of commercial or ego-soothing benefit? Lets share thoughts about and to grow the positives?

I wonder of Facebook will handle this but after the ageing question looked at this population that I specifically imagined rings to be associated. Not much joy from that except to get an idea of pupping frequency. I enjoyed seeing these pictures again and will post all my Tulista. This lot are the famous Lemoenpoort variant – check that glabrous one!

No nice pictures here but this is at Mowers (an abandoned railway gangers cottage). Road construction between Worcester and Robertson led to destruction of part of a large population of T. pumila. For some incomprehensible reason these plants have been burdened with a species name. It is difficult even to say it is a discrete population because walking either to Worcester or Robertson, or even north or south, you will find plants for many miles. It would be a flip of a coin to say they were the same or different.

Some greenish plants on old river gravel and across the valley on high shales all shades of chocolate in abundance. How I regret not filling my camera with those incredible plants if pictures can ever do justice to them.

Two plants from very near Lemoenpoort demonstrating the 8 leaf-tier formation of the Fabonacci spiral series. It is usually between 8 and 13 for the species. The single plant is from only the one population I know of where the plants are much smaller and darker, and the leaf tiers are in the 3-5 tier range. In fact when I first came across them I felt a deja vu kind of moment that I was east of Oudtshoorn looking at H. scabra (tuberculata).

Martin Scott is very knowledgeable and fascinated with Tulista and these are from near where he grew up near Witteberg way to the NE of the distribution of pumila. See one plant dividing. Warwick made some great comment on plant ageing and dividing among other points, but Facebook format tends to tuck such away.

My interest in Haworthia is totally a biocentric one. An anthropocentric interest, touched by the way we are propagandised by science, religion and economics is bound to cause differences and disagreements. These pumila pictures were taken northeast of Stormsvlei but south of the Riviersonderend river. Oh why in this digital era was I so stingy with camera capacity? Could shed tears for so much lost in the celluloid one.

What makes Haworthia including Tulista, so fascinating is the extraordinary local variability. Many more pictures would be needed to display that at this place east of Worcester at the base of the Langeberg.

These were photographed virtually within Barrydale and I am sure have now been destroyed. I believe they are a bit further east too, but interestingly there are fertile hybrids with Astroloba corrugata (muricata) about 14km east of Barrydale that suggests a more solid presence.

A last picture of pumila taken at Worcester but at probably the most western point of occurrence. The plants were blood red in summer 1970 during a prolonged drought. I went back perhaps 20 years later hoping for the same dramatic colour! Next I am going to go through my T. minima pictures with a great deal of withhold. Someone has found it necessary to change the name to minor. On of those totally unnecessary names justified by virtue signaling a Rafinesque like intent to create an impression. I fear that plant taxonomy is facing a meltdown and the aloids are leading the subject into (further?) disrepute. I will keep touching on “why”.

Fascinating stuff this business about plant classification. Up to about 2000AD there were 4 major systems. The Engler system for Europe, The Bentham Hooker one for England (and Commonwealth), the Tahktajan one for USSR and Cronquist for USA. Since DNA technology, we have the Angiosperm Phyllogeny Group (APG) trying to generate consensus on the basis of a better understanding of evolutionary history. A cursory inquiry suggests there are 13 members/contributors – 2 Swedish, 4 British (Kew 3), 1 Australian, 5 USA and 1 Finnish. On the practical fun side this is Tulista minima at the western most end of the Potberg.

Apart from the four major classification systems there comes the question of national and Independent herbaria and their choices and decisions in respect of whatever system they adopt. For the Aloids, the principals seem to be IOS (International Organisation for Succulent Plant Study), Kew, SANBI (South African National Biodiversity Institute) and Tokyo Institute for Evolutionary Biology? Recently it was said that IOS had overturned the Tulista recognition, but that is not so. IOS never recognised it in the first place. SANBI has adopted a system of classification based on DNA evidence that does not seem to take into adequate account the aloids north of the Limpopo. It has omitted some SA species too. These pics are Tulista minima from the northern Potberg area locality and I have several of those.

IOS is based on he Zurich botanical garden and has 97 private members. Kew has still to make its presence felt and then of course there is the role of herbaria world wide and a ton of individual interest from all directions. always new ideas and new brooms on the scene. Always a potential Rafinesque, or Resende or an icon of intellectual virtue finding his or her way in the strange world of taxonomy. It is simply a constantly changing scene. It is not science. It is an expression of our view of the world and how we interact with each other? More from the Potberg flats.

The Aloids are in a bit of a limbo and really caught up in the idiosyncrasy, egocentricity and autocracy of plant taxonomy, nomenclatural rules, institutions and individuals. It is NOT good and the future looks bleak. The path of Haworthia since Linnaeus is laughable if it was not so tragic. I know the last 80 or so years very well and particularly, and intimately, the last 40 years. It has all been a sheer left brain debacle. So it is nice to look at this large T. minima from far SW Heidelberg.

Of course T. minima is characteristically blue-green in colour but in the Bontebok Park we get these. It has got something to do with T. marginata but exactly what. Maybe it has something to do with T. pumila too, or is it even a bigger system that included T. kingiana. There are or were two other populations that are/were similar. It is all well and good tinkering with the nomenclature and using fancy left-brain technology to arrive at some overworked mechanistic outcome, but does it help?

There really is just an unimaginable range of variation even in Tulista where classification does not seem to be controversial. But when he eventually see what Essie feels he has learned from 40 years of field observation, you might begin to ask yourself if our species concepts are inline with reality. I agree with him.

These minima are from west of Riversdale. Taxonomy is practiced much as if one dances the Charleston or Jerusalema while navigating between shops at a mall. Much display but not necessary!

Small Tulista minima from SE Heidelberg. That is Euphorbia tridentata in the first pic with a weird distribution. Its home is actually NE Cape. But I am going to go on about the aberrations of classification and “science”. Sometimes (how often is that?) oddballs/taxonomists/amateurs/novices/experts/wannabees describe or described specimens and not species.

People from all walks of life have been messing with the Aloid genera for the last 70 years to the point that recent events are simply continued absurdity. Despite the stand-off between IOS and SANBI, a new single species (unispecies, monotypic???) genus has been published for Aloe suzannae. To what end? Just what purpose does this serve? It appears that DNA sequencing does not even play a role. Aloe suzannae is reported to be in the Aloidendron clade any way. What about all the species that are not yet covered by the published sequencing data – as suspect as I am sure it is. When will someone who actually knows the aloids recognize that Aloe striatula is wildly odd, as is Aloe haworthioides. Has sequencing really successfully buried all the other questionable genus/species combinations? What does the statement mean concerning the Aloid phyllogram that reads that there does not appear to be evidence of reticulate construction? Trust these Tulista minima will bring some solace or else take an aspirin. Is this ordinary minima in a grassy fire-threatened habitat?

A real life experience. A top ranked taxonomist revising a genus examined the material available to him and identified and described two species. Species X with the four character states A1 B1 C1 D1 and species Y with the alternate states A2 B2 C2 D2. But I was dealing with new living material and none of the plants I saw matched eg. A1 B2 C2 D1, or any other combination PLUS new ones of something like A3 or C5. I wonder if there ever were even the types that reflected the taxonomist’s decision! This has happened to me many times. Not only that – as I learned about Haworthia, I had to change my opinion many times too. More to come about that. But this is a Tulista population very near to T. marginata by about 100m – clearly there are ‘hybrids’. But remember the other green T. minima from about 1km away in the Bontebok Park! I will show more Bontebok park populations and you will begin to see that maybe Tulista is a single system.

Please note that I have had many of those disconcerting “real life experiences”. There is an astounding level of naiveté among budding taxonomists who really over-reach themselves in the process of expounding what amounts to factually limited real knowledge. Often done in the most complex language. As an entomologist and then as “assistant” at the Karoo Garden, my prime function was identification of field material, visits to the herbarium and contact with “experts”. If I appear cynical, it is with very good reason. These Tulista minima are from several populations along the northern base of the Potberg.

What really distressed me was the problem Larry Leach had in an unending argument with Kew about the validity of a Euphorbia name that Welwitsch had collected and referred to in his notes as “candelabrum”. Kew argued vehemently that Welwitsch had never intended to name it “candelabrum”. What was their motive in resisting the name? Why? They did not like Welwitsch because he had been funded to collect by Kew and ended up selling his collection to another herbarium! Larry’s obituary unkindly reads that he wasted valuable time arguing the issue about the name. This just demonstrates that taxonomic nomenclatural issues are often decided like modern day law suits. The lawyers lose nothing however the cases pan out, and the more appeals the better. Not to forget the food for the obituarist! Let’s go back to another Bontebok park population. Do you think it is possible to categorically state that these are T. minima and not a small pumila?

I want to post some more relevant and recent stuff so want to get past Tulista and touch on this nonsense of multivariate analysis and the obfuscation of “science”. This is another Bontebok park population of minima. Very pumila looking but they are smallish and I have many flower pictures to show the pinkish coloration rather than the olivaceous of pumila.

Nearly a meter across. T. minima from S Bredasdorp. Long ago before all the cultivation I came across a similar population of proliferating plants on sandy flats close to the lower Breede River. They were much smaller and compacted together in such enormous clumps.

We struggled a bit to find these minima/marginata plants a few days ago … W. Heidelberg … next is what used to be.

A second (B) population further north also of these minima/marginata confoundments.

This is A2 when there were many plants at A1. These plants are extremely sensitive to animal hooves!!

Still further north we find plants like this that we regard as T. marginata – spectacular?

East of Bonnievale are these slender, erect leaved marginata and there are ‘hybrids’ with both pumila and minima – and again the hybrids are not interspersed with supposed parents.

Here are the “pumilaXmarginata” hybrids (as a small, local population)…

I should have had a lot more photos of this population i.e. marginataXminima? Neither supposed parent was in the immediate vicinity by several hundred yards. The hybrids were fertile and I grew a lot from seed. There is another population of minima westwards by a few km and there too are this time with pumila. Again the hybrids are in a discrete small area away from either supposed parent.

This locality is in Smith’ collecting record. NE Bredasdorp. I struggled to find it until I realised that it was one of the many fatalities of a vandalistic attitude of road builders. In the 1970s they simply ignored any environmental concern and ravaged any surface stone as well. I am not sure if either minima or marginata were in near vicinity but this is the mix and named (in my opinion needlessly) as a discrete species viz mortonii. A few plants do still survive there.

Retusa Travels

Retusa Travels was first published as a series of Facebook posts from May and June 2022.

Here are pictures of Kweekkraal Haworthia mirabilis MBB8064 – some plants can be deemed to be atrofuscoid, others magnificoid and even heidelbergensoid. Kweekkraal has come up before in my exposé of “H. mutica var. nigra” and at this stage of my journey I conclude that the “nigra” issue was just one small issue showing that there is a single system that unites retusa (including turgida) and mirabilis.

More MBB8064 … Quite a lot on the mind with a mind boggling amount of new material. Talking to Lawrence and Essie about utilizing Haworthiaupdates.org more intensely as publication is very time consuming and also very limiting in the sense of illustration that tells most of the story. Kweekkraal is some 10km west of Riversdale and the area is the focus of my first article on the H. mutica ‘nigra’ anomaly. There is so much feeling about the disclosure of locality that I think is counterproductive. These plants are just as much under stress from porcupines, grazing animals, other farming activity, while there are clear laws in place about trespass and plant removal. My number for this collection is MBB8064, a new and recent ‘location’ and it was incredibly difficult to find the plants at all. People need to switch to camera capture, and I will post more photographs of this site to demonstrate variability. One could say H. mirabilis “atrofuscoid”. i.e. many if the plants are reminiscent of what was described as H. atrofusca. The name can be kept and used in that sense while technically the actual Latin binomial refers to a very complex and large SYSTEM with many variables. So a formal name is really just a matter of convention and agreement.

Essie brought this population of mirabiloids to my attention. It is also Kweekkraal as part of Kiewietsvlakte that figures in my first essay on “How to understand H. mutica var. nigra“. H. floribunda occurs as discrete and also lost in the milieu. H. retusa is present. H. mirabilis occurs as the atrofuscoid form in the east, a magnificoid cocktail in the middle and heidelbergensoid to the west. Kweekkraal is a Guest Farm and there is a large area of conserved Renosterveld. My reference number for these is MBB6085.

More of MBB6085 at Kweekkraal west of Riversdale. A mirabilis potpourri. Do look at the shrinkage of the plants. This is at the end of the dry season and there would normally have been rain by now. Plants can look very different as the seasons come and go. Who can tell me the difference between a low-end intellectual enjoying plants in the field, and a top-end intellectual in a white coat sitting at the controls of the Large Hadron Collider trying to manipulate dark matter? There is a lot in this question!!

The last of MBB8065 – a point of interest is porcupine activity. I have seen this again and again and also where the damage is extensive. I reported it to conservation management who replied quoting a “scientific” report on porcupine activity at Nieuwoudtville where there is very little threat of land transforming to agriculture, The conclusion is that Porcupines are an essential part of the ecosytem. When they do become a nuisance in urban areas they are trapped and moved to conservation areas. Well I agree. Under natural conditions porcupines and nature go together. But enter man and his greedy and destructive ways; the situation is different.

This retusa (retusoid) is another of Essie’s finds and is very close to the splendens (mirabiloid) localities which would suggest that they might be discrete systems. This is not the only situation that does this, but looking at all our known populations, we still conclude that it is the nature of the beast as it is drawn and quartered by nature! Incidentally, we struggled to find plants where many had been reported. Why? Immense porcupine activity. There used to be floribunda at this place too. Since degraded by agricultural activity and is destined to be totally cultivated in the near future.

With Essie near a retusa pilgrimage spot. Essie has explored so much of this area and here he is indicating what he has seen and where. The plants here were in bad shape compared with some previous visits and porcupines have also been very busy. Sometimes chewing off the leaves and the plants survive, but more often grubbing them out.

Re those silver flecks. I feel constrained to post this here and now. There has been and is so much ill-feeling and misunderstanding in Haworthia classification that I have been accused of initiating while also bearing the brunt of it. Here is a quote from a great Haworthia personality who finds me unbearable …”the silvery flecks in the leaves of H. jakubii suggest connection to H. magnifica and H. pygmaea.” That is interesting because in the photograph MBB7818 Windsor 2, there are clearly silver flecks that this exceptional man denies it is determination to prove me wrong about the species status of H. jakubii. I do not find them in any of my magnifica (mirabilis) pictures from the type locality or in MBB6651 from nearby, but I do find them in H. retusa. I would be so grateful if this was shared around and brought to the attention of all Haworthia enthusiasts. Essie has been caught up in the mess too and I am enlisting his help to solve the problem. We have together written two articles in the process and are stuck in the third seemingly because of the wish to avoid further injury.

Essie showed me these retusa at the Riversdale town edge. Municipalities universally feel it is necessary to have a great swath of vandalistic dumping to signify their dominance over nature and make human presence known. So its a bit painful to walk on the margins and bid what will surely be a goodbye. But do also watch for flecks and see if you can decide how you would describe H. retusa.

Daphne Bayer exploring.

More retusa from west of Riversdale. This species also has that enormous range of variation that makes classification so awkward and contentious. Some of those quirky leaf shapes and tips occur in H. mutica too.

I could quite happily go back to Riversdale for another 20 pictures of these captivating plants. I was on about “white flecks”. Us mad taxonomists are always looking for characters by which to make outrageous claims, but some of these things like flecks and windows can be pretty behavioral beyond just variables in a population.

The significance of the flecks is that it is critical in relation to the recognition of jakubii as a species in view of the arguments put forward by proponents. The argument is made that jakubii has these flecks whereas my collection at Windsor a little further is supposed to not have them. Further argument is made that my photographs are just fleeting images and bear no relation to close observation in cultivation. So I can show that these are plants at Windsor that do have flecks, not only that, they appear in more species where this antagonistic proponent implies their absence. Both Essie and I simply observe that jakubi is in the continuum of H. mirabilis, and the fleck argument is void.

This is a photo of a pygmaea from Humor. These plants were named argenteo-maculosa because of the silver markings. There are actually two kinds. The narrow sharply defined and the more diffuse larger markings along the veins. The former could be oxalate crystals and the latter aggregates of chloroplast filled cells. It should be understood that there are more than 20 features in leaf anatomy that some scientists feel could actually be used alone to identify down to species level. My experience and opinion is that it only leads to very emphatic opinions than are not always true. In fact generally quite false. An example is my experience where I was called in as a practical field observer to support results obtained for surface studies where atomic absorption photography had been used to make big decisions. Why was that necessary at all?

More retusa – it is quite instructive to examine windowing in the individual plants. It would be very difficult to quantify. Flecking either as chlorenchema aggregates or oxalate crystals if that is what they are – is relatively absent.

This diagram of interconnectedness and continuity has its roots in a first draft drawn in 1966 when it was pretty clear that there were clinal gradients from population to population of Haworthia. This has been demonstrated time and time again. Essie has independently come to exactly the same conclusion from the same careful and considered exploration and observation. All these elements on the map are somehow connected and are best understood in this context. This has nothing to do with simply providing labels and names for collectors – it is about providing a basis for a deeper appreciation of the plants and their origins. In fact, of nature itself.

The blue is the mirabiloids, pink the turgidoids, red retusoids, orange rossouwoids, red crosses muticoids. There are species names in the map. Just remember the lines connect discrete localised populations. Ideally each populations should be “identified” and individually mapped.

What now? I think I am going to carry on with retusa as the red-line drawn on that map. It is totally implicated in turgida as a complementary element. The two are really different in respect of habitat occurrence and degree of proliferation AND not always! I have had these all published and I have posted before, but I want to write with new emphasis supported by what Essie has to say. He is posting on his page… Haworthia veld memories and we are busy writing together as a team. Together we can make this map more significant. These are Essie’s “close east of Riversdale” retusa.

Another West of Riversdale population with pictures taken by Warwick. Take note that populations differ from one another just as the plants within them do too.

These are about the most southerly of retusa.

I have often taken a pilgrimage to this site S Riversdale and the population always seems a bit different. I posted a few pictures of plants from here a few days ago taken May this year. The present pictures were taken 1st Aug 2012. Bud are just pushing. Wait until you can compare these with pictures taken 2008 and 2020. Habitat pics should be time-stamped to take into account the shifting dynamics of field populations. Perhaps it would be better to say that some research ought to be undertaken to explore population level changes. Just note that many of these populations are not somewhere you just walk out the back door to see. With modern photography I would suggest a minimum of 20 good photographs for any one habitat – which means take about 50! The underlying truth is that taxonomy has to move away from tinkering with names, to considering the impact of time and conditions.

Sept 2008 S Riversdale. Plants in flower and looking great?

January 2020 and the plants colouring up in the sun.

This is another of Essie’s discoveries and particularly interesting because it highlights the reality of the relationship between the recognised species. I regard this population as Mirabiloid retusa because over the whole Southern Cape there is ample evidence that despite gross differences in flowering time of the two “species” involved, they do “hybridise”, but more significantly that they are part and parcel of one system, which calls for a better definition of what the word “species” actually means.

This is close to east Riversdale where there used to be four populations of this plant. The southern one was gobbled up by cultivation and was the origin of the name “fouchei” that was an erect leaved non-proliferating form of H. retusa. Semblances of that form occur in the next population that is some 300m north …

A few hundred meters north is another very well known population H. retusa “geraldii”. When I first saw these plants ca 1971 the plants were in huge flat clusters over 400mm diam. With time the clusters broke up and were far less impressive. Retusa is normally a fairly non-proliferating element.

There is direct continuity between turgida and retusa. They do not “mix”. I also think now that it is incredibly more complex and that we only have the two groups … the very dark green late-summer flowering mirabilis and the light green/yellow spring flowering turgida. These two match and mix to produce a host of variants of which the main are mutica, pygmaea and emelyae. I do not know quite how herbacea/rerticulata/maculata/pubescens come out in the west.

I wrote in 1981 …”The species problem very seldom arises where species grow together, but rather from the difficulty in associating populations strung out in isolated populations.” This in itself is a rather weak explanation. Here I have just posted two H. retusa populations but there is a third population that I ascribe to H. mirabilis that others say is H. asperula. Curiously I took two sets of pictures for that because I found there to be two sub-populations closely adjacent … this is one of them…

In contrast. About 30m away in vegetation unprotected from grazing animals for the past 80+years, thus impacted vegetation and skeletal soil, is this bunch of variables.

Mirabilis Travels Introduction

Mirabilis Travels was a series of Haworthia shucked Facebook posts in 2022.

There is not much that I can now usefully do so I am going to switch attention to H. mirabilis. If any one species exemplifies the complexity of the genus it must be this. When I first started organising field information I ran headlong into host of names with very little to relate them to in the field. Now I recognize that it is that mirabilis forms a dark coloured summer flowering half of a single system that extends from southwest of Bredasdorp all the way to east of Mossel Bay, while also extending into the Little Karoo. In no particular order, here is the “var. notabilis” from east of Worcester. My great regret is that I have no digital record of a great stretch of exploration prior to about 2002.

Wheeeee!! It really stretches the imagination. Just randomly diving in was a bad idea. So I add fuel to the fire by showing these in the no-man’s-land between “systems”. This mirabilis is west of Swellendam and both this and the previous “notabilis” cast light (or shadow) on the relation of the retusa (turgida)/mirabilis system and that of herbacea/reticulata/maculata/pubescens of the Worcester/Robertson area. The near presence of a floribunda variant is added value. One could almost argue for a restatement of the situation where mirabilis and turgida are established as generic names with many species (population variants) attached. Notice how I interchange turgida and retusa – I explain that the taxonomic system trips itself with its own rules. Priority means that retusa holds sway over turgida, when in fact it is more probable that turgida is the genetically and historically dominant.


I have over 200 folders of mirabilis localities just since 2000 and no picture records for the 30 preceding years. Much posted for this species in HaworthiaUpdates.org, but nothing can help dismiss the unbelief that these could all be one species. Here is just a blind selection of 7 pictures to show how improbable it all seems … imagine another few hundred variants!

2. Meiringii

These are mirabilis “meiringii” from east of Bonnievale. There must be something significant in the similarity to H. herbacea? Bear in mind we are dealing with many populations strung out over the landscape in what Essie and I recognise as continuities. But, mirabilis is not continuous with herbacea. Flower and flowering time as well as geographical location and continuity with many other populations show “mirabiloid”.

A little further east = still “meiringii”

… some more from a bit further east before the more traditional fewer leaved forms occur …

This is what I wanted to post – more meiringii from almost at Bonnievale. Curiously this is also the only place in the Robertson Karoo where Aloe glauca occurs. It is salutory to compare the taxonomy of Aloe vs. Haworthia because the scales are quite different. However, A. glauca also has a range of variants that never seem to have been examined. I can’t imagine what the classification of either will look like after another 50 years of this limited one-man-band style of taxonomy. Meiringii does seem to have some degree of independence but just how much and guessed at by whom? It diffuses to a spineless spotted form westward, which in turn diffuses to three different versions following compass direction.

3. Mirabiloid

How does one get to grips with H. mirabilis? If you take all the available names for this lot of dark coloured February – March flowerers and match them up against the 400 or so known populations, you are going to have to explain why most of the mirabiloid populations are in a nameless limbo? This is a population from North Ashton is quite out of character for that area.

These are at the southern entrance to Cogmanskloof and fairly similar to plants northwards as well as to the general ensemble of mirabilis in the Worcester/Robertson Karoo, but I am having fun putting together a general perspective of H. mirabilis. It was not even considered known in the field until I started messing with the classification. That is another story I do not want to see unfold! So far there are 44 names that cover the story of the mirabiloid to help explain why so few have any idea of what the name mirabilis conveys.

I have had a bit of fun in taking all the names relevant to H. mirabilis to show what a taxonomic nightmare it has been and will remain. Those 50 extant names only represent about 75 populations at the most. In the majority of cases they do not cover even the variants in the designated populations. So not only do we have another possible 200 or more populations to consider, but also the variation within them.

This is about the most westerly record for mirabilis in an almost ecological desert. There is clear pattern in how the populations relate to one another, with just enough exceptions to overturn expectation.

4. Napier

There are about 10 known populations of this type of mirabilis between Caledon and Napier that gave rise to the varieties of triebneriana viz. napierenis, turgida and multituberculata. Grasping what the situation is with this system of species would go a long way to resolving the differences of opinion about the entire genus.

As we move eastward to north of Napier the plants change a bit as they get smaller. It is worth looking for that floribunda-like leaf tip!

At Napier itself the plants are smaller. Just keep in mind we are considering just what constitutes this species mirabilis. It is a questioning of trying to reconcile about 300 or more populations across a very wide area AND accommodating them each and everyone in a rational system. Remember history gives us about 50 names many of which are alternate species names. How do we make sense of this? So I am taking what pictures I have and local area by local area presenting the problem as it is – not what I imagine it to be.

5. Badia

Badia is a particularly striking variant of mirabilis and occurs as several populations along a 3km stretch of a recent geological surface west of Napier. Forming a part of the southwest border of distribution of the species.

These are badia from the western population – rougher surfaces.

6. Mundula

Bredasdorp is only 15km east of Napier and has its own odd set of mirabilis. Mundula is from southwest of the town. In 1969 when I first saw the plants, there were large clumps of 30 to 40 rosettes, highly coloured. Over the years the clumps collapsed, the surrounding vegetation aged and the plants were scattered about and either single or in much smaller clumps.

8. Rossouwii

About 6km northeast of Bredasdorp March 2006. This is what mirabilis is really about – a whole host of populations from Riviersonderend in the west to well east of Riversdale in the east. From inland of Montagu in the north to a report of these plants on the cliffs shore west of Infanta. Small unglamorous, cryptic, often non-descript, infinitely variable, beneath interest to many wannabees. This particular population of rossouwii is a tiny remnant that survived a massive road building program in the 1970s that vandalised surface rock. Do check those leaf ends!

H. rossouwii occurs as 3 populations WNW Bredasdorp with no close presence of mirabilis. Then it jumps a huge distance to SSW Heidelberg where it also never has close association with mirabilis. But at Bredasdorp, east of rossouwii there are a heap of small mirabilis populations that include some curious variants between stubby leaved and elongated leaved, as also degrees of spination. This population is directly north of Bredasdorp.

These plants are not very photogenic as they avoid light and are invariably tucked away in tight spots. Each plant seems to have its own microhabitat and will look different if grown anywhere else. So it is always difficult to fully grasp or describe some kind of common denominator for a population. In cultivation they grow out of character too.

9. Heidelbergensis

While generally presenting the things that constitute mirabilis, I have mentioned rossouwii. It seems to be generally assumed that the species are real discrete things that can be fitted into some sort of classification structure. This is definitely not true. At one stage I thought there might be a species heidelbergensis, and consequently named a population north of Bredasdorp as its var minor. So let us look at the issue. Before I get to that particular population, look at these plants to the west …

When my hard drive crashed I was just going to start the mirabilis heidelbergensis conundrum. It is not an area I like at all because there is no clarity as already indicated. So here is a population close to the origin of Smith’s “heidelbergensis” – but these are big plants and I file them as retusa/turgida when they could better be turgida/mirabilis or retusa/mirabilis. It is so easy to generate opinion and comment when there is not much information available. So I made a selection of photographs of 20 or so populations to indicate what has to be incorporated in a classification – somehow or other! Often pictures are poor and then I only have since about 2004. Another problem is seasonal difference. One is not always lucky to get to a population when the plants are filled out and actively growing, while they are not the same year by year either. Much is made of growing and observing these plants in cultivation. Bananas! All very well for ten, twenty even 100 populations. But impractical for hundreds.

About 800m west of the type locality for “heidelbergensis” and 400m N of the previous post of large plants, these are small and roughly equate to ‘heidelbergensis’. Here the plants are turgid and greenish coloured as opposed to the much darker and shriveled in the dry season or spell. Being dark coloured they are invariably well sheltered under other vegetation.


H. mirabilis is generally a dark coloured plant. Thus mirabiloids are dark coloured vs. the retusoids(turgidoids) that are lighter. Of course in shade or cool conditions the plants green up. Here is a “light coloured” mirabilis from Greyton to just demonstrate that making any statement to distinguish mirabiloid from retusoid is difficult. My conclusion is indeed, because they are the same species! “To avoid confusion” I must also add that when I say that plants that look different are the same species, some critics get highly apoplectic? This is the problem my posts are addressing and there is not much evidence available to me that the message gets received or is welcome. Perhaps the latter sentiment is most true because I find it uncomfortable myself.

Just east of Heidelberg is another population of these small plants that fall in the ambit of what actually is heidelbergensis. Growing in think moss and lichen it is a slightly different habitat. These pictures are of plants in cultivation.

These are a few km S Heidelberg. Now I am enthralled by what my own pictures are telling me. I write to find out and establish what my observations mean! I just deeply wish I had prepared sufficiently for the end stage when the pieces fall into place. What is happening here in floribunda is included in the compromise. Nearby we have a retusa var. ‘nigra‘ for which the story has been told. ‘Heidelbergensis’ is for me a deep insight into the connection between the mirabiloids and the retusoids (turgidaoids). Colour is a significant character, so is the bend back of the leaves as well as the rounded leaf-tip. A thing to note is that “retuse” as originally used was to mean “bent back like a thumb” and NOT the modern usage for a flat leaf in which the margins grow ahead of the leaf point. So erect vs retuse leaves is also significant, but in Haworthia it is not a species determinant and more than colour or rounded leaf-tip OR even flowering time. These pictures help to demonstrate this fact. If just one population did this we could perhaps dismiss the issue as hybridization. And this is my greatest regret. I just do not have enough or good enough pictures taken at all seasons and of more variants, or for all the populations I have been so privileged to enjoy. Because i am going to be succeeded by “taxonomists” intent on imposing their own opinions for the sake of effect rather than reality?


Eastwards the ‘heidelbergensis’ form extends half way to Riversdale where it merges with the atrofuscoid/magnificoid versions of mirabilis. But this population is further south down the Duiwenhoks growing in close proximity to retusa ‘nigra’ and the putative parent for that viz H. turgida. There is now so much evidence for the unifying of formal names and especially considering Essie’s exhaustive exploration and opinion, independently validating what I have to say.

Still further down the Duiwenhoks are these two populations for which there are sadly not enough pictures to remotely demonstrate the variation. What is perhaps worth noting is leaf shape, colour and translucency. Still further down the river is a major conundrum in the form of a population of dark unvariegated could be variegate. This is a problem of distribution because along the coast there are no haworthia between Stilbay in the east and the Duiwenhoks and then to Infanta in the east, apart from paradoxa. It is a major issue as there is no continuum as for the inland strip.

A few kilometers SE Heidelberg is this population of ‘heidelbergensis’ and I will follow with a further one nearby that demonstrates the transitions from retusoid to mirabiloid. This population is nevertheless already mirabiloid as is evident by the darker colours and the summer flowering time. In the last post I mentioned an aberrant population of presumably variegata. While trying to explain the complexity of species generally, I have avoided some serious issues with populations that simply do not fit anywhere, while they, only in their oddity. can they be claimed to fit a rational species definition. I hope to come to those.

Moving about 1km further SE is this population that also demonstrates variability and the nature of population differences. This population is more magnificoid in that there is a tendency for retusion of the leaf, for more textured surfaces and for less translucency coupled with more blackish-green coloration.

Before continuing with ‘heidelbergensis‘ just look at these magnifcoid/atrofuscoids from west of Riversdale where there is no turgida. Different habitats and also it is the retusoid version that does occur just here but infrequently.

These non-descript mirabiloids also from west of Riversdale can only roughly be distinguished from what I refer to as Heidelbergensoids. They occur from well east of Riversdale all the way to north of the Potberg, Bredasdorp, Riviersonderend, into the Worcester/Robertson area to south of Worcester, to Ashton, Robertson and then into the Little Karoo at Montagu and Barrydale. If you scrambled different quantities of varied colour thick paints on a board and scratched them around with a stick, heidelbergensis would appear as a thin web within the middle of the outcome.

In my Revision I recognised 4 varieties of heidelbergensis and this was really because I did not know what to do with these troublesome variables. Twenty five years later and with a lot more information, I can only consider a sensible explanation to be that they are part and parcel of a single system viz. H. mirabilis (and of course intimately linked to retusa/turgida). Here are two pictures of my first var. toonensis from SW Heidelberg. Unfortunately only two cultivated specimens and no idea of variation, but there is a lot better to come. Essie may have more because these are down the Slangriver that he knows well. What is so valuable about Essies conclusions is that he has arrived there on the basis of a wholly different set of populations and individual plants – and without herbarium support or influence.

10. Turgida

Also southeast of Heidelberg is this population of turgida just to demonstrate how variable it also is. Here it has erect narrow leaves and shows the colouration of the retusoids. Haworthia turgida occupies about 75% of the area that mirabilis does but they never actually share habitat. I remind you that I do not like the nomenclatural system and where it occurs that the name retusa would be required to be preferred over turgida, when in fact retusa is a lesser variant of turgida.

11. Slangriver

These are a bit west of the Slangriver and taken in July during the wet season. So the plants are green, turgid and about as much translucency as they can show. It occurs to me now that I always thought of words rather than pictures, but in this digital age this is about how many pictures needed and of better quality.

Essie Esterhuizen commented “It was in this area where I realised in the early 90’s that I don’t have the ability to sensibly distinguish between the then maraisii, magnifica, mirabilis and heidelbergensis.”

To which Bruce Bayer replied. “Well Essie, its not a question of your ability. It simply cannot be done. Those names just existed because of the history of the subject and even now there is surely some doubt that you and I are stating the fact that a true classification does not meet the perceptions or needs of enthusiasts and collectors. Their need are real and it is time amateur journals started assuming some responsibility for a way of using names that is useful, while being accurate.”

This is also west of the Slangriver and not just making a mockery of any attempt to subdivide H. mirabilis into heidelbergensis, atrofusca, jakubii, magnifica, maraisii, shuldteniana and many more, you can clearly see that floribunda is involved too. I do not know of any floribunda southwest of Heidelberg except the evidence that appears in these small mirabiloids that this is where it has gone. Soumen Aditya is a good observer and it will be nice to know how his opinions of the names that are developing.

Soumen Aditya replies “Sir, true…one population and its species concepts no value at this moment. Horrible variations each single species and definitely link each others. Floribunda is a simple variation under many mirabiloid included maraisii. Thank you.”

This is a few km downstream on the Slangriver from toonensis and a great opportunity for someone to explore the prediction that more populations will exhibit the similarities that follow geographical order. There are some unique plants in this post and the Slangriver is no stranger to oddity. It is not easily accessible. It follows a strange path – first southward and then westwards to flow into the Breede River between Malgas and Infanta where there are many mirabilis and some turgida variants.

Some way to the northwest and the plants seem to have lost translucency while the leaves are shorter stouter and perhaps darker green. But there are MANY populations of this ilk varying in one direction or another, supporting the truth of Essie’s comment that categorizing them by any of the more relevant names is just not possible.

12. Zones

While considering what to post next on Mirabilis Travels I realized that I have to break mirabilis up and organize my pictures better. There are practically speaking 12 geographical zones and I only have pictures for about 200 populations of at least 400 I have visited that fall into the category I recognize as H. mirabilis. I cannot believe that anyone can dispute the statement that these populations and the internal variation form a continuum and any taxonomic subdivision will be purely artificial. In fact I go further and state that the species H. turgida, retusa, pygmaea, mutica and emelyae are more probably than not, included. When the involvement of H. floribunda with it’s associations, and then the probability that H. emelyae has further connections are considered as well, the enormity and complexity of the situation can be imagined. But here are pictures from two further populations to go with the previous SW Heidelberg.

Each of those mirabilis zones has something going on pointing to the complexity I refer to, but right now I am posting about the heidelbergensis mirage. This is the Swellendam zone and the population one from S Buffeljagsrivier. The home of groenewaldii, that is a variant that for me puts the whole retusa/mirabilis complexity into perspective.

This, further south from Buffeljagsriver. The plants were not in good shape and this is just one of the sad facts of life. It raises the issue of how to decide what it is. But the situation is that there is no clear idea of just what constitutes heidelbergensis. West of Buffeljagsrivier there are 6 known populations of mirabilis that I cannot state why exactly they are the same. There is an obvious similarity in respect of size, colour and flower but confoundment with the floribunda/variegata variant as well. Can it be understood and can it be explained? So lets rather continue southwards down the Breede River for the moment.

Further south along the Breede River is a “farm” where we found 8 populations of mirabilis and I will just show plants from 3 localities to show how they vary. See the floribunda influence too – floribunda as a recognizable population is absent south of the Bontebok Park but reappears as such down along the western end of the Potberg. But remember we are actually on a heidelbergensis mission to show that it is fiction just as magnifica and maraisii (and others) are.

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13. Multifolia

Eureka – have I got it? For years I have been pondering the problem of how and why the form of rossouwii appears in mirabilis ’heidelbergensis’, mirabilis ’sublineata’, and emelyae ‘multifolia’. Essie has independently indicated that he observes this too. I should actually be travelling, in my series of posts about heidelbergensis, from west to east to show this. Sadly I do not have pictures of the heidelbergensis I saw further east of Heidelberg, nor any of Essie’s for the eastern Hessequas area. DNA does not create form. It only provides the materials to do so. Form depends on the creative power of the metaphysical element (causal mind – consciousness) of each life form? I will put a question mark there again, because this is a serious issue we need to contemplate. The photos I add are multifolia variants from 4 populations from Little Karoo. The last one touches on Warwick asking why I fingered a single variant from S of “heidelbergesis var. toonensis” along the Slangriver.

14. Rossouwii to Bromberg

So we are still south of Buffeljags along the Breede River. Looking for heidelbergensis evidence. Not much in this lot but I did also interject with a post referring to rossouwii. Very interesting because now one has to consider all populations, related species AND their distribution. Because this is a whole and species cannot be summarily treated like cherries or plums and picked by whim and fancy. Haworthia rossouwii is very odd because it occurs at two centers about 30-40km apart and then appears to reoccur in a smaller form just south of Heidelberg.

Bit of a dilemma. Rather than go south I will for the moment go west from Swellendam towards Stormsvlei. The end will shock as it really demonstrates the impossibility of arriving at a solution on the mechanistic platform.

The predictability of a working hypothesis? Essie sent me this lovely picture of a plant from the mountains between Heidelberg and the Little Karoo. But like me Essie has only just awaken to the fact that you cannot get people to believe what you say unless you can give them a greater visual experience than you had yourself? Essie has a LOT to tell us. This is a link between the turgidoides (i.e. retusa and mirabilis together therein) and H. emelyae. It will eventually be recognized to also include rossouwii?

A little further west in the Luiperdsberg area west of Swellendam.

Now between Luiperdsberg and Bromberg on the way to Stormsvlei. There is not a smooth transition and a huge number of populations nevertheless supports the contention that this is a single system viz H. mirabilis.

I don’t have pictures of mirabilis on the south side of the Bromberg so these are from north. Also I am going to skip going still further north into the Worcerster Robertson Karoo and beyond the Riviersonderend River. I wrote all about mirabilis in Updates about 2006 and don’t have pictures for the complexity in that area and westwards, which would help to ease the oddity of transformation.

Tony Brook asked if mirabilis is the oldest name. Bruce answered “A very uncomfortable question. I have no doubt that it is atrovirens – but it is a tricky path and I am no fan of the nomenclatural code or the fancy footwork that goes with it. Goes back to Stearn and typification of old names.”

These Bromberg variants will surely excite the splitters but really when one sees so many of the populations from McGregor to Swellendam as well as to the north; and even the ecological semi-desert to the south, you have no option but to abandon an impossible point of view. Unfortunately the population of ‘heidelbergensis v. scabra’ (i.e. before my even conservative view was proved hopeless) at Leeurivier east of Drew, was ploughed out despite the non-arable nature of the shale. But there are pictures in Revisited and also in early volume of Updates re: H. mirabilis.

Authoritarianism is a real problem in present society and this comes up in plant classification too. So it is not compulsory to see these plants as H. mirabilis too. It is just my personal opinion. The plants on the north slopes of the Bromberg are in sandstone and these now are along the Riviersonderend River in manganic tertiary deposit. This is very unusual habitat. Taken late February as they flower in the heat of summer.

These were taken 3 years earlier and In January when cooler wetter conditions prevail. There are four microhabitats in a very small area.

Now this is a real freak for which I have no explanation. A single small population in a weird habitat on the south side of the Bromberg. Sandstone slabs and very wet. About 200m away is shale and also what one expect of H. mirabilis. But this flowers in November and has in-turning leaves that one finds in, say herbacea. Although there are individuals, as you can see, in which the leaves are erect-spreading. The flowers are nothing unusual except the buds are flattened top to bottom and the ends also turned down slightly like-in mirabiloids. So it throws everything out of the window or into disarray. It does not alone meet the requirements of a species and there is zero indication of what its real affinities may be. My deeper feelings are that it mirrors a dark side of (at least Haworthia) classification and that the real species is the entire “genus”. For the present I have speculated, as I have suggested in this series, that floribunda/variegata/chlorocantha are intimately connected to the turgidoids, and suspect that rossouwii is as well. I have treated this H. eliseae as a variant of rossouwii, as I have done with ‘calcarea’, ‘petrophila’ and my own ‘heidelbergensis var. minor”. Let me add something. Classification is not some thing confined to part of the group so that you can cherry-pick where and to what you want to apply a name. A classification has to be ALL inclusive and the specimens remaining in the incertae folders are only there because they are too poor to permit any decision.


About 200m away is H. mirabilis.

15. Down the Breede to the Potberg

The myth of heidelbergensis has dissipated to the west and north and morphed into others, so let us go back down the Breede River and see what mirabilis holds for us in that direction. There’s a gap that I haven’t explored, the 20km stretch of the Breede River as it twists and turns south of Swellendam. The next two populations are south of Malgas, east of the river.

Here are more from the east bank of the Breede River south of Malgas. Note that the populations on the west bank all along the northern lower reaches of the Potberg are incredibly complex with names like joleneae, bobii confounded with atrofusca, floribunda, variegata, turgida, and mutica.

As an after thought … did anyone see this? Some how or other these two forms of H. eliseae came about, and I suspect it may have something to do with Rupert Sheldrake’s suggestion of morphic fields. Is this an overlap of the Worcester/Robertson herbaceoids and the more southerly mirabioids? I settled for a rossouwii/mirabilis interaction that is not entirely convincing. These two pictures were taken at the same time and place in one small population

I am not sure how to deal with the west bank populations. Essentially I am talking about H. mirabilis and have been exploring the heidelbergensis phenomenon. On the lower west bank of the Breede River we have a long conjunction of shale and sandstone mixed with tertiary deposits. The vegetation is either fynbos or renosterveld and there are vast differences over short distances. H. turgida in its best form is present (along the river itself) and so is the floribunda/variegata element. The result is two contrasting versions of mirabiloids. I will post the 6 or so turgidoid influenced populations first (fynbos) and then the more dark mirabiloids (Renosterveld) after. Note that there are no absolutes. On the decaying shales where there is kaolinite, we could be back west of Riversdale. I will start with what deviously and/or distastefully, acquired the name ‘bobii’.

This is just a few km west of bobii and again a link in a chain that makes a mockery of populations as species. I don’t think the Potberg is well explored at all. While this is on the eastern and north facing flank, there are two other populations known on the southern slopes travelling westwards.

On the southern face of the Potberg a few km further west and well off the beaten track …

Again travelling west along the south side of the Potberg is a similar population of plants. Unfortunately I only have two pictures of plants in cultivation that are not very useful except to show how growing conditions affect appearances. We arrived at site late afternoon in rainy weather. It was a long hike through thick brush and the light was also too poor for film photography.

Leaving the Potberg, returning to bobii at Infanta and then searching along the lower slopes between the Potberg and the Breede River. Just in what respect and by what margin do the next three populations differ from the previous 4? This was March 2006.

It is a bit of overdose because these are pictures of the same population as previously but taken a year later in September.

Accession

 

Accession Name Map1 Map Location
decipiens var cyanea 3222 DC Trakas Kuilen
mucronata var inconfluens 3321 AC 8km W Ladismith
mirabilis var consanguinea 3419 BA Die Galg
limifolia var limifolia 2631 BD Mbuluzi gorge, Blue Jay ranch
mucronata var inconfluens 3321 CA 9km SW Ladismith
heidelberg var scabra 3420 AD W Kathoek
cooperi var pilifera 3227 BD Fort Murray Bridge
reticulata var reticulata 3319 DB Buitenstekloof
parksiana 3422 AA Botteliersberg
pygmaea var argenteomaculosa 3421 BB Cooper Siding
mutica var mutica 3419 DB Klipdale
pumila 3319 CD Lemoenpoort
sordida var sordida 3325 AC Brakfontein
pulchella var pulchella 3320 CA Nougaspoort
reticulata var reticulata 3319 DC Dublin
herbacea var paynei 3319 DD 1km S McGregor
pygmaea var argenteomaculosa 3421 BB Humor
marginata 3420 AC Adoonskop
pulchella var pulchella 3320 CB SW. Anysberg
pumila 3319 BC Osplaas, DeDoorns
kingiana 3422 AA Little Brak, Barswell
semiviva 3120 CD SE Middelpos
marumiana var marumiana 3124 CC 32km E Murraysburg
mutica var mutica 3420 AB Crodini
1404a cooperi var tenera 3325 CB Groendal Dam
2469a nigra var diversifolia 3221 CB W Merweville
4168b glauca var glauca 3324 AA 2km E Humefield
6512a maraisii var maraisii 3420 AA SE Drew
6584a glaucaXviscosa 3324 AD Waaipoort
6584a glaucaXviscosa 3324 AB NE Waaipoort
KG100/74 blackburniae var blackburniae 3321 AC W Ladismith
KG100/740 blackburniae var blackburniae 3321 DB Warmbaths
KG107/74 magnifica var magnifica 3420 BB SW Heidelberg
KG109/74 mucronata var morrisiae 3322 CA Volmoed
KG111/72 bayeri 3322 CA S Oudtshoorn
KG114/72 emelyae var comptoniana 3323 AD Georgida
KG115/71 arachnoidea var nigricans 3321 DB Warmwaterbron
KG118/70 arachnoidea var arachnoidea 3319 DA Effata
KG118/71 emelyae var major 3321 CC Muiskraal
KG119/72 scabra var morrisiae 3322 AD Schoemanspoort
KG120/71 mucronata var morrisiae 3322 CB Hazenjacht
KG125/73 arachnoidea var nigricans 3321 DA S Calitzdorp
KG126/72 arachnoidea var nigricans 3321 DA S Calitzdorp
KG126/72 arachnoidea var nigricans 3320 DD E Lemoenshoek
KG138/72 emelyae var major 3321 CD Sandkraal
KG140/72 decipiens var decipiens 3323 BB Constantia
KG146/72 bayeri 3322 CB Doringkloof
KG151/72 mucronata var morrisiae 3322 CB Kamanassie Dam
KG155/73 fasciata 3325 CB Hillwacht
KG160/72 arachnoidea var setata 3321 CB S Ladismith
KG160/72 arachnoidea var setata 3322 BC Meiringspoort
KG163/70 maraisii var maraisii 3319 DD S Goudmyn Bridge
KG166/70 herbacea var herbacea 3319 CB Brandwacht
KG166/71 arachnoidea var nigricans 3321 CC Springfontein
KG167/71 arachnoidea var nigricans 3321 CC W Springfontein
KG175/70 reticulata var reticulata 3319 DC Rooiberg
KG193/73 cooperi var gordoniana 3324 DD 5km N Hankey
KG202/70 magnifica var atrofusca 3421 AA Spitzkop, W Riversdale
KG218/70 herbacea var herbacea 3319 DC Mowers
KG224/70 maraisii var meiringii 3320 CC E Bonnievale
KG231/70 decipiens var decipiens 3323 AD Skerpkop, E Willowmore
KG240/72 turgida var turgida 3420 BB N Heidelberg
KG241/72 turgida var turgida 3420 BB N Heidelberg
KG257/77 emelyae var emelyae 3321 DC SE Vanwyksdorp
KG26/70 mirabilis var triebneriana 342O AA Stormsvlei
KG28/70 mirabilis var triebneriana 3419 AB N Uitkyk
KG2/71 maraisii var meiringii 3320 CC W Bonnievale
KG30/70 mirabilis var triebneriana 3419 BA near Greyton
KG310/70 bolusii var blackbeardiana 3227 AC SW Cathcart
KG311/7 magnifica var acuminata 3421 BD N Gouritzmond
KG311/70 bolusii var blackbeardiana 3227 AA Imvani
KG312/70 bolusii var blackbeardiana 3126 DD Bowes’ farm
KG315/70 decipiens var minor 3324 AA Campherpoort
KG31/70 mirabilis var triebneriana 3419 BA near Greyton
KG323/70 mucronata var morrisiae 3321 BC Bergplaas
KG324/71 reticulata var subregularis 3319 DA Boskloof
KG326/71 maraisii var maraisii 3420 AA Rondeheuwel
KG327/71 mutica var mutica 3420 AC N Kykoedie
KG329/70 herbacea var herbacea 3319 DD Koningsrivier
KG329/72 nortieri var nortieri 3118 DC Die Kom
KG32/70 mirabilis var beukmannii 3419 BA Skuitsberg
KG336/70 gracilis var gracilis 3326 AB Hellspoort
KG345/71 maraisii var maraisii 3319 DD 9km W Robertson
KG34/70 turgida var longibracteata 3420 AC 20km N Bredasdorp
KG35/70 maraisii var maraisii 3420 AC 19km N Bredasdorp
KG36/70 heidelberg var minor 3420 CA Rooivlei
KG36/71 arachnoidea var arachnoidea 3319 BD Tonnel Stn.
KG382/70 cooperi var pilifera 3326 DC Brigadoon
KG383/70 cooperi var pilifera 3326 DA Peninsula
KG388/71 arachnoidea var setata 3321 DA E Vanwyksdorp
KG392/70 cooperi var cooperi 3227 AC W Cathcart
KG393/70 bolusii var blackbeardiana 3227 AC SW Cathcart
KG402/70 springbokvlakensis 3324 BD Springbokvlakte
KG411/75 chloracantha var chloracantha 3321 DD N Herbertsdale
KG436/75 decipiens var cyanea 3323 AD Georgida
KG441/75 arachnoidea var nigricans 3321 BC N Calitzdorp
KG46/70 maraisii var maraisii 3319 DD 18km Robertson to Bonnievale
KG47/72 gracilis var tenera 3326 BA Plutosvale
KG47/72a gracilis var tenera 3326 BA Plutosvale
KG535/71 arachnoidea var setata 3321 CB W Vanwyksdorp
KG567/71 mucronata var inconfluens 3321 AC Dwarsrivier
KG568/71 mucronata var morrisiae 3321 CB W Vanwyksdorp
KG573/51 arachnoidea var nigricans 3321 CB Ladismith to Vanwyksdorp
KG593/71 arachnoidea var setata 3321 CB 14km E Ladismith
KG623/69 mutica var mutica 3420 AC Soesriver
KG627/69 retusa 3421 AB Near Riversdale, Grootvlei
KG628/69 mirabilis var badia 3419 BD Napier
KG630/69 maraisii var maraisii 3319 DD 9km W Robertson
KG631/69 turgida var suberecta 3422 AA Brandwacht
KG640/69 arachnoidea var arachnoidea 3319 CB 8km NE. Worcester
KG662/69 reticulata var reticulata 3319 DA Keeromskloof
KG669/69 maculata var maculata 3319 CB Brandvlei Dam
KG681/69 mirabilis var triebneriana 3419 BD Mierkraal, Napier
KG682/69 mirabilis var triebneriana 3419 AB Uitkyk
KG688/69 maraisii var maraisii 3319 DD SW Robertson
KG692/69 mirabilis var triebneriana 3419 BA Near Genadendal
KG695/69 herbacea var paynei 3319 DD McGregor
KG6/72 cooperi var leightonii 3327 BA Kaysers Beach
KG77/71 mucronata var rycroftiana 3321 DD 23km E Ladismith
KG7/71 maraisii var meiringii 3320 CC W Bonnievale
KG806/70 longiana SE Hankey
KG83/71 magnifica var magnifica 3421 AA S Riversdale at beacon
KG83/77 mutica var mutica 3420 AC Kykoedie
KG85/72 mutica var mutica 3420 AC Badjieskraal
KG89/76 reticulata var hurlingii 3320 CC Goudmyn
KG90/76 reticulata var reticulata 3319 DC Rooiberg
KG92/71 magnifica var magnifica 3421 AB E Riversdale
KG93/71 variegata var variegata 3421 AB Hoekraal
KG94/71 turgida var longibracteata 3421 AC Brakfontein
KG98/71 chloracantha var subglauca 3422 AA E Great Brak
KG9/71 maraisii var meiringii 3320 CC W Bonnievale
? marginata 3320 CC NE Ashton
112 limifolia var gigantea 2731 DA Nongoma
121 maraisii var notabilis 3319 DD Vinkrivier
153 arachnoidea var arachnoidea 3319 DB Buitenstekloof
154 minima var poellnitziana 3320 CC W Drew, Swellendam
155 lockwoodii 3320 BB Floriskraal
157 parksiana 3422 AA Dumbie Dykes
158 floribunda var floribunda 3420 BB 6km N Heidelberg
159 mucronata var inconfluens 3321 BC Die Berg
160 reticulata var reticulata 3319 DC Ribbokkop
161 herbacea var herbacea 3319 DC Ribbokkop
163 pulchella var pulchella 3320 AC Avondrust, Touws River
163 pubescens var pubescens 3319 CB Sandberg Hills
164 maculata var maculata 3319 CB Brandvlei Dam
166 serrata 3420 BA Oudekraalkop, Heidelberg
167 venosa ssp woolleyi 3324 BD Kleinpoort
168 venosa ssp venosa 3420 AB Swellendam
169 attenuata
170 glabrata
171 arachnoidea var nigricans 3322 AC Schoemanspoort
171 mucronata var mucronata 3322 AD Schoemanspoort
172 marumiana var marumiana 3226 AB Spring Valley
173 bolusii var bolusii 3225 CA NE Pearston
174 marginata 3421 AB S Heidelberg
175 monticola var monticola 3322 DD Molen River
886 fasciata 3325 CB Spring Range
887 fasciata 3325 CC 8km E Hankey
1119 maculata var maculata 3319 CB Audensberg Peak
1120 maculata var maculata 3319 DA S Sandhills
1128 pubescens var livida 3319 CD S Lemoenpoort
1145 maculata var maculata 3319 CD Moddergat
1194 coarctata var coarctata 3326 DB Fairfax
1208 maraisii var notabilis 3319 DD Wolfkloof, Robertson
1209 maraisii var notabilis 3319 DD Wolfkloof, Robertson
1210 maraisii var maraisii 3319 DC Trappieskraalkloof
1211 maraisii var maraisii 3319 DD W Robertson
1213 maraisii var maraisii 3420 AA 7km N Stormsvlei
1214 maraisii var meiringii 3320 CC W Bonnievale
1215 maraisii var maraisii 3319 DC Langkloof
1216 maraisii var maraisii 3319 DD Goudmyn
1217 maraisii var meiringii 3320 CC W Bonnievale
1218 maraisii var meiringii 3320 CC W Bonnievale
1219 maraisii var maraisii 3320 CC N Drew
1220 maraisii var maraisii 3319 DD Klaasvoogds
1221 maraisii var maraisii 3420 AD Juliusfontein
1221 maraisii var maraisii 3319 DD Skurweberg
1222 maraisii var maraisii 3319 DD S McGregor
1351 coarctata var adelaidensis 3326 AB Hellspoort
1352 coarctata var coarctata 3326 AB Hellspoort
1354 coarctata var adelaidensis 3326 BC NW Grahamstown
1355 coarctata var adelaidensis 3326 AA Willowfountain
1357 coarctata var coarctata 3326 BC Brooklands
1358 coarctata var coarctata 3326 BC Manley Flats
1359 coarctata var coarctata 3326 DA Kowie
1360 coarctata var coarctata 3326 BC Vaalvlei
1361 coarctata var coarctata 3326 DA S. Vaalvlei
1363 coarctata var coarctata 3326 CB Salem to Alexandria
1364 coarctata var coarctata 3326 DA Hopewell, 20km W Port Alfred
1366 coarctata var coarctata 3326 DA Hopewell, 20km W Port Alfred
1367 coarctata var coarctata 3326 BA Rooidrift
1368 coarctata var coarctata 3326 BA E Plutosvale
1370 coarctata var coarctata 3326 CA Woodbury
1371 coarctata var coarctata 3325 DB Orlando
1372 coarctata var coarctata 3326 AD Yellowwoods
1381 reinwardtii 3326 BD Frazer’s Camp
1391 reinwardtii 3327 AA 3km W Peddie
1438 reticulata var reticulata 3319 DC Rooikleigat
1539 reticulata var reticulata 3319 DC S Gemsbokkop
1543 reticulata var reticulata 3319 DD Wolfkloof, Robertson
1558 emelyae var multifolia 3321 CC Muiskraal
1558 emelyae var multifolia 3321 CC Springfontein, Riversdale
1615 arachnoidea var nigricans 3321 CA SW Ladismith
1616 arachnoidea var nigricans 3321 DB Oudtshoorn to Calitzdorp
1617 arachnoidea var nigricans 3321 CA Adamskraal
1618 mucronata var habdomadis 3321 AC Dwarsrivier
1619 arachnoidea var namaquensis 3219 DD Ceres Karoo
1620 cymbiformis var cymbiformis 3326 BA E Fort Brown
1621 cooperi var leightonii 3327 BA SW Paynes Hill
1621 mucronata var morrisiae 3321 DB Warmbron
1622 cooperi var pilifera 3326 AA Willowfountain
1623 cooperi var pilifera 3227 DA St Johns Drift
1624 arachnoidea var nigricans 3321 CA W Ladismith
1628 mucronata var inconfluens 3321 CA 4.5km S Ladismith
1652 arachnoidea var namaquensis 2817 CD Kliphoogte
1661 venosa ssp tessellata 2817 CD Kouefontein
1674 arachnoidea var namaquensis 2917 AB Karrachabpoort
1698 heidelberg var toonensis 3420 BB Matjestoon
1700 heidelberg var scabra 3420 AB Leeurivier
1701 mucronata var rycroftiana 3321 DC Van Wyksdorp to Herbertsdale
1702 zantneriana var minor 3323 BB near Miller Station
1703 maraisii var maraisii 3319 DD W Robertson
1704 arachnoidea var nigricans 3320 DD Warmwaterberg
1706 cymbiformis var setulifera 3228 BA S Mooiplaas
1707 maraisii var maraisii 3319 DD Muiskraalkop
1708 maraisii var maraisii 3320 CC N Ashton
1963 arachnoidea var scabrispina 3320 CA Nougaspoort
1986 mucronata var inconfluens 3320 DA Anysberg Pass
1995 herbacea var herbacea 3319 CD W Doornrivier
1996 herbacea var herbacea 3319 CD Lemoenpoort
1997 herbacea var herbacea 3319 DC Wansbek
2022 bolusii var bolusii 3224 BA Graaff-Reinet
2024 cooperi var cooperi 3225 CB Bruintjieshoogte
2038 marumiana var marumiana 3126 BD N Tarkastad
2052 angustifolia var baylissii 3325 BC Oudekraal
2070 decipiens var pringlei 3224 DB Ebenezer
2071 bolusii var bolusii 3224 DD Lootskloof
2072 bolusii var bolusii 3224 BA Graaff-Reinet
2074 gracilis var viridis 3324 AA Campherpoort
2076 decipiens var minor 3324 AA Grootriver, Mara
2083 decipiens var cyanea 3323 CA N Uniondale
2086 arachnoidea var aranea 3322 DC Ganskraal
2093 pulchella var pulchella 3320 AB SE Konstabel Stat.
2099 mucronata var morrisiae 3322 CA 8km S Oudtshoorn
2105 arachnoidea var scabrispina 3320 BA N Baviaans Stn.
2117 viscosa 3220 DD 43km N Laingsburg
2124 arachnoidea var scabrispina 3220 DC 35km N Laingsburg
2131 viscosa 3220 DD 43km N Laingsburg
2177 maraisii var maraisii 3320 CC Goedverwacht
2187 reticulata var reticulata 3319 DD Wolfkloof, Robertson
2241 pygmaea var pygmaea 3422 AA Great Brak
2271 maraisii var maraisii 3319 DD Houtbaai Kloof
2287 pygmaea var pygmaea 3422 AA W Great Brak
2289 pygmaea var pygmaea 3422 AA Dumbie Dykes
2311 floribunda var dentata 3421 BA Wydersrivier
2347 marumiana var marumiana 3224 AB Valley of Desolation
2350 nigra var nigra 3224 AB St Olives
2356 marumiana var marumiana 3124 DC Aasvoelkrans
2372 venosa ssp tessellata 3222 BC E Molteno Pass
2373 marumiana var marumiana 3222 BA Molteno Pass
2377 decipiens var cyanea 3221 CB W Merweville
2380 bolusii var bolusii 3124 CC W Graaff-Reinet
2385 venosa ssp tessellata 3223 AA Nelspoort
2389 bolusii var bolusii 3123 DD E Murraysberg
2406 semiviva 3222 BC S Beaufort West
2418 arachnoidea var nigricans 3321 CD W Vanwyksdorp
2419 arachnoidea var nigricans 3321 CD S Vanwyksdorp
2420 turgida var longibracteata 3420 AB 5km SW. Swellendam
2421 herbacea var herbacea 3319 CB W Worcester
2422 herbacea var herbacea 3319 CB SE Brandvlei Dam
2423 magnifica var acuminata 3421 BD N Gouritzmond
2424 wittebergensis 3320 BC SW Laingsburg
2425 lockwoodii 3320 BB Ezelsfontein
2453 marumiana var archeri 3221 CA Langberg
2453 mirabilis var beukmannii 3419 BA Skuitsberg
2453 mirabilis var triebneriana 3419 BA Dagbreek
2526 mutica var mutica 3420 AA Rietfontein
2533 turgida var longibracteata 3420 BC Diepkloof, S Malgas
2547 heidelberg var scabra 3420 AC Brakfontein
2550 heidelberg var heidelbergensis 3420 BB E Heidelberg
2551 variegata var modesta 3420 AD SW Kathoek
2556 heidelberg var scabra 3420 AD Beyersdal
2564 variegata var hemicrypta 3420 BC NE lower slopes of Potberg
2574 arachnoidea var namaquensis 3219 DC Skitterykloof
2579 herbacea var lupula 3319 CD Wolfkloof, Villiersdorp
2582 pulchella var pulchella 3319 BD W Touws River
2591 maculata var maculata 3319 CB NE Brandvlei Dam
2665 magnifica var atrofusca 3421 AA Droerivier
2670 venosa ssp venosa 3420 BC Malgas
2672 turgida var longibracteata 3421 AC Duiwenhoksriver
2697 herbacea var herbacea 3319 DC Keerweerder, Jonaskop
2716 arachnoidea var nigricans 3321 CA Winkelplaas
3363 monticola var monticola 3323 AD Georgida
3375 gracilis var viridis 3324 BC Dorschfontein
3384 monticola var monticola 3323 CA E. Uniondale
3398 arachnoidea var namaquensis 3119 AB Koringberg
3423 arachnoidea var namaquensis 3119 BC Beeswater
3439 floribunda var dentata 3420 BA Bontebok Park
3453 venosa ssp venosa 3420 AB Bontebok Park
3586 chloracantha var denticulifera 3421 BB Cooper Siding
3586 chloracantha var denticulifera 3421 BD 3km N Gouritzmond
3612 arachnoidea var nigricans 3320 AB Jagerskraal
3620 marumiana var viridis 3322 AC S Prince Albert
3637 nortieri var nortieri 3118 DC W Doornriver
3906 nortieri var pehlemanniae 3320 BB 5km SW Laingsburg
4160 bolusii var bolusii 3323 BB NE Fullarton
4179 glauca var glauca 3224 CD Fairview
4180 decipiens var cyanea 3224 CD Fairview, W Jansenville
4198 gracilis var viridis 3325 AC Brakfontein
4294 nigra var diversifolia 3222 AC Wolwehoek
4320 arachnoidea var namaquensis 3220 DA Verlatenkloof
4404 cooperi var gordoniana 3323 CA Uniondale Poort
4428 blackburniae var blackburniae 3321 DA Rooiberg
4429 blackburniae var blackburniae 3321 DA Assegaaibos
4430 herbacea var paynei 3319 DD Olifantsdoorn, McGregor
4432 minima var minima 3420 AB Bontebok Park
4437 maraisii var maraisii 3319 DD W McGregor
4438 mucronata var inconfluens 3320 DA Jakkalsfontein
4439 herbacea var herbacea 3319 CD N Lemoenpoort
4440 decipiens var cyanea 3322 BC E Klaarstroom
4450 cooperi var pilifera 3326 AD NW Salem
4460 cooperi var pilifera 3227 DC Brigadoon
4461 maculata var intermedia 3319 DC Buitenstekloof
4462 arachnoidea var aranea 3322 AC S Cango Caves
4462 arachnoidea var scabrispina 3320 CA 20km W Ladismith
4471 turgida var suberecta 3421 BA Weltevrede
4473 mutica var mutica 3420 AC Soesriver
4474 cooperi var gordoniana 3324 DD NE Hankey
4475 cooperi var isabellae 3324 DD NE Hankey
4476 cooperi var isabellae 3324 DD 2km E Hankey
4476 turgida var suberecta 3421 BA Draaihoek, Albertinia
4477 turgida var suberecta 3421 BA E Valsch River Bridge
4478 turgida var suberecta 3421 BB Gouritz River
4479 mutica var mutica 3419 BB 10km E Riviersonderend
4479 turgida var longibracteata 3421 AB Kafferkuils Bridge
4499 arachnoidea var nigricans 3322 AC N Oudtshoorn
4549 cymbiformis var cymbiformis 3325 BC Kranspoort, W. Patterson
4575 arachnoidea var nigricans 3320 DD W Warmwaterberg
4642 mirabilis var triebneriana 3419 BD Skietpad
4647 arachnoidea var aranea 3322 AD Raubenheimer Dam
4648 cymbiformis var ramosa 3327 AB Wooldridge
4648 cymbiformis var cymbiformis 3327 AB W Woolridge, Peddie
4649 cymbiformis var reddii 3226 BB Klipplaat Dam
4650 cymbiformis var obtusa 3326 AC S Alicedale
4651 cymbiformis var obtusa 3226 DA Blinkwater
4652 cymbiformis var cymbiformis 3326 BB Ballinafad
4653 cymbiformis var obtusa 3325 BB
4654 cymbiformis var cymbiformis 3327 AC Kapp-Fish confluence
4655 cymbiformis var cymbiformis 3226 DC Sulphur Baths
4656 arachnoidea var nigricans 3322 CA Volmoed
4657 decipiens var cyanea 3324 BD Hanekam
4659 reticulata var hurlingii 3320 CC Goudmyn
4665 reticulata var attenuata 3320 CC 5km SE Bonnievale
4665 reticulata var reticulata 3319 DD Wolfkloof, Robertson
4667 venosa ssp tessellata 3225 BA Halesowen
4668 venosa ssp granulata 3320 AC Touwsriver, Avondrust
4669 venosa ssp granulata 3219 DC Skitterykloof
4671 arachnoidea var nigricans 3320 DC E Barrydale
4677 venosa ssp tessellata 3123 CD Nuwerus
4677 heidelberg var scabra 3420 AA Kliphoogte
4901 marginata 3420 BA Koppies
4902 serrata 3420 BA Koppies
4941 sordida var sordida 3325 AD SE Kirkwood
5086 semiviva 3120 DB S Williston
5096 semiviva 3120 DC S Williston
5101 heidelberg var scabra 3420 AD Haarwegskloof
5157 decipiens var decipiens 3322 AB Kleinsleutelftn, Prince Alb.
5182 decipiens var decipiens 3322 AA W Prince Albert
5209 marumiana var marumiana 3221 DD Tierberg
5225 scabra var scabra 3322 AC Cango
5226 scabra var scabra 3322 AC Cango
5261 decipiens var decipiens 3322 CA S Prince Albert
5296 mucronata var inconfluens 3320 CB Touwsfontein
5296 mucronata var morrisiae 3320 CB Touwsfontein
5750 venosa ssp granulata 3220 CC Bantamsfontein
5753 arachnoidea var setata 3321 DA E Vanwyksdorp
6486 arachnoidea var scabrispina SW Laingsburg
6488 venosa ssp tessellata 3120 Beaufort West Dam
6488 semiviva 3120 BC Beaufort West dam
6490 venosa ssp tessellata Karoo National Park
6495 nigra var nigra E Merweville
6496 nigra var nigra W Merweville
6497 decipiens var cyanea 15km Merweville to Koup
6498 marumiana var marumiana 3222 BA Molteno Pass
6501 semiviva Langberg
6502 nigra var nigra Langberg
6505 nortieri var nortieri 3219 AC N Dwarsrivier
6505 nortieri var nortieri 3219 AC N Dwarsrivier
6506 nortieri var nortieri 3219 AC Wolkberg Cracks
6506 nortieri var nortieri 3219 AD Dwarsrivier
6509 heidelberg var scabra 3320 CC W Bonnievale
6509 maraisii var maraisii 3320 CC 0.25km W Bonnievale
6510 maraisii var maraisii 3320 CC 5km W Bonnievale
6511 maraisii var maraisii 3319 DD Goudmyn W River
6512 mutica var mutica 3420 AA SE Drew
6512 mutica 3420 AA SE Drew
6513 mirabilis var triebneriana 3420 BD 3km SW Swellendam
6514 maculata var intermedia 3319 DD Buitenstekloof
6515 reticulata var acuminata 3320 CC Aurora, Bonnievale
6516 maraisii var meiringii 3319 DD Rooiberg, Goudmyn
6518 maraisii var maraisii 3319 DD Lower N Slopes, Rooiberg, Goudmyn
6519 maraisii var maraisii 3319 DD Agter Vink
6520 maraisii var maraisii 3319 DD Die Nekkie, W Robertson
6521 maraisii var maraisii 3420 AA N Stormsvlei
6522 arachnoidea var arachnoidea 3319 DD Buitenstekloof
6523 venosa ssp venosa 3420 AB Swellendam
6524 limifolia var limifolia Komatipoort
6525 rossouwii var calcarea 3420 AD De Hoop
6527 variegata var variegata 3420 AB NW Swellendam
6528 floribunda var floribunda 3420 AB SW Swellendam
6533 maraisii var maraisii 3420 AB N Napky
6534 maraisii var maraisii 3420 AA SE Drew
6535 reticulata var reticulata 3320 CC N slopes Rooiberg Goudmyn
6536 mutica var mutica 3420 AB S Napky Witklipkop
6537 mirabilis var triebneriana 3419 BC NE Goudini
6538 mirabilis var triebneriana 3419 BD 10km W Napier
6539 maraisii var maraisii 3420 AD Tarentaal
6540 minima var minima 3420 AD Tarentaal
6541 heidelberg var scabra 3420 AD NE Kathoek
6542 variegata var modesta 3420 BC E Potberg residence
6544 heidelbergensis 3420 BC NN boundary Potberg reserve
6545 heidelbergensis 3420 BC inside N Boundary Potberg
6546 heidelberg var scabra 3420 AD NW Kathoek entrance
6548 Bosfontein
6551 floribundaXmagnifica NE Cooper siding
6552 fasciata 3324 DD NE Zuurbron
6553 cooperi var gordoniana 3324 DD 5km N Zuurbron
6554 cooperi var gordoniana 3324 DD 8km N. Zuurbron
6555 cooperi var isabellae 3325 CC Longmore Forest
6556 bolusii var pringlei 3225 DD NW Rippon Stn.
6557 cooperi var pilifera 3325 BB NW Ripppon stn
6558 cooperi var dielsiana 3225 DB N Eastpoort
6559 cooperi var dielsiana 3225 DB SE Eastpoort
6560 cooperi var dielsiana 3225 DB S Eastpoort
6561 bolusii var pringlei 3225 DB Baviaanskranz, Patryshoogte
6562 cymbiformis var obtusa 3226 CD Kagasmond
6563 cooperi var cooperi 3226 CD Koonap Bridge
6564 cooperi var dielsiana 3226 CB Chancery Hall
6565 cooperi var dielsiana 3225 DA 5km W Somerset East
6566 bolusii var bolusii 3225 AC NE Ashbourne
6567 cymbiformis var reddii 3226 BB Klipplaat Dam
6568 nigra var nigra 3225 BD Klipplaat Dam
6569 bolusii var blackbeardiana 3226 BD Waterdown Dam
6570 bolusii var blackbeardiana 3226 BD S Estrelle
6571 bolusii var blackbeardiana 3227 AB Turnstream
6572 cymbiformis var reddii 3227 AB Turnstream, SE Queenstown
6573 cymbiformis var setulifera 3227 AB Highclere
6574 nigra var nigra 3224 DA 5km ENE Kendrew
6575 nigra var nigra 3226 CB Adelaide
6580 decipiens var virella 3224 DC Meerlust
6581 decipiens var virella 3224 DC Welgelegen
6582 decipiens var virella 3324 AB SW Mt Steward
6583 decipiens var virella 3324 AB NW Waaipoort
6584 glauca var herrei 3324 AD Waaipoort
6585 zantneriana 3324 AB NE Waaipoort
6585 zantneriana var zantneriana 3324 AD Waaipoort
6586 glauca var herrei 3324 BC Zeekoeisnek
6587 decipiens var minor 3324 BC NW Die Bordjie
6588 sordida var lavranii 3324 BC NE Die Bordjie, Baroe
6589 gracilis var viridis 3324 BC NE Dorschfontein
6591 cooperi var pilifera 3226 DC S The Tower, Ft Beaufort
6592 cooperi var dielsiana 3226 DC W Fort Beaufort
6593 cymbiformis var cymbiformis 3226 DC W Sulphur Baths ne Tower
6596 kingiana 3322 CC Moeras River
6597 cooperi var gordoniana 3424 BB Jeffrey’s Bay
6598 glauca var glauca 3325 CB Bauerskraal
6599 cooperi var pilifera 3325 CB Bauerskraal, Uitenhage
6600 cooperi var viridis 3325 AC N Perdepoort
6601 arachnoidea var aranea 3322 CC Moeras River
6602 cooperi var pilifera 3326 BC Glen Craig, NE Gstwn
6603 cooperi var gracilis 3326 BA 7km NE Grahamstown
6604 arachnoidea var xiphiophylla 3325 DC Coega
6608 arachnoidea var setata 3322 CB N Dysselsdorp
6609 truncata 3322 CB N Dysseldorp
6610 arachnoidea var setata 3324 AC N Steytlerville
6611 sordida var sordida 3325 DA Soutkloof
6612 aristata 3325 DA Soutkloof
6613 sordida var sordida 3325 BC Bluecliff Stn
6614 cooperi var gracilis 3326 AB Hellspoort
6615 glauca var glauca 3325 AC Paardepoort
6616 decipiens var xiphiophylla 3325 CA Bauerskraal
6618 decipiens var minor 3325 AC Sapkamma/Perdepoort river
6619 decipiens var minor 3325 AC Sapkamma/Perdepoort S slope
6620 decipiens var minor 3325 AC Sapkamma entrance
6621 outeniquensis 3322 CC Moerasriver
6622 pumila 3319 DA Mowers
6623 pumila 3319 DC W Rooiberg
6624 minima var poellnitziana 3320 CC W Drew, Swellendam
6625 maraisii var maraisii 3320 CC Sandrift Drew
6626 heidelberg var scabra 3320 CC N Sanddrift
6627 marginata 3320 CC N Sanddrift
6628 minima var poellnitziana 3320 CC Sanddrift
6629 marginataXminima 3320 CC E Sanddrift
6630 minima var minima 3419 DB W Moddervlei, Elim
6631 mirabilis var mirabilis 3419 DB Mierkraal
6632 rossouwii var calcarea 3420 CA Renosterfontein, Karsriver
6633 marginata 3420 AC Adoonskop
6634 marginata 3420 AC Adoonskop
6635 mirabilis var badia 3419 BD Napier
6636 mirabilis var triebneriana 3419 BA S Greyton
6637 minima var minima 3419 DB Mierkraal
6638 maraisii var maraisii 3420 AC Rooivlei, Bredasdorp
6639 mirabilis var sublineata 3420 CA S Bredasdorp
6640 maraisii var maraisii 3420 AC Adoonskop
6641 mutica var mutica 3420 AC Hasiesdrift
6642 pumila 3419 DD Agtervinkrivier
6643 mirabilis var triebneriana 3419 BD 15km W Fairfield
6644 mirabilis var triebneriana 3420 BD 3km SW Swellendam
6645 pumila 3319 CB Worcester airfield
6646 maraisii var maraisii 3319 DD N Macgregor
6647 maraisii var maraisii 3319 DD AgterVink
6648 maraisii var maraisii 3319 DD SW Robertson (Oliva)
6649 arachnoidea var setata 3320 CC 4km E Montagu
6650 mutica var nitida 3420 BB Diepkloof, 1km SE Heidelberg
6651 magnifica var magnifica 3421 AA 3km S Riversdale
6653 emelyae var emelyae 3321 CD N Sandkraal
6654 emelyae var emelyae 3321 CD N Sandkraal
6655 emelyae var emelyae 3321 CD N Sandkraal
6658 arachnoidea var aranea 3321 CD E. Sandkraal
6659 emelyae var emelyae 3321 CD SE Vanwyksdorp to Herbertsdale
6660 emelyae var multifolia 3321 CC W Muiskraal farmhouse
6661 arachnoidea var nigricans 3321 CC W Muiskraal farmhouse
6662 magnifica var atrofusca 3421 AA Kweekkraal road
6663 magnifica var magnifica 3420 BB 2-3km SW Heidelberg
6666 magnifica var magnifica 3420 BA S Tradouw Pass
6667 maraisii var maraisii 3320 DC SW Barrydale
6668 maraisii var maraisii 3320 CC N. Ashton
6670 maraisii var maraisii 3319 DD 12km W Robertson to le Chasseur
6672 maraisii var maraisii 3319 DD Koningsriver turn-off
6673 maraisii var maraisii 3319 DD McGregor Nature Reserve
6674 pumila 3319 DD McGregor Nature Reserve
6676 maraisii var meiringii 3320 CC E Goudmyn
6678 maraisii var maraisii 3319 DD Grootrivier, SW Goudmyn
6680 herbacea var paynei 3319 DD Koningsriverberg
6681 maraisii var maraisii 3319 DD Koningriver Dam
6682 maraisii var maraisii 3319 DD 2km N Koningriver Dam
6683 maraisii var notabilis 3319 DD N Klaasvoogds
6684 reticulata var attenuata 3320 CC E Dankbaar, SE Bonnievale
6685 mirabilis var diversicolor 3320 BB Olifantsdoornkloof
6686 mirabilis/maraisii 3320 BB Olifantsdoornkloof
6687 maraisii var maraisii 3319 DD SE McGregor
6688 herbacea var herbacea 3319 DA Mowers
6690 arachnoidea/mucronata 3320 CA Watervalkloof, Rietvlei 2
6691 maraisii var maraisii 3319 DC NW Boschfontein
6692 reticulata var reticulata 3319 DC NW Boschfontein
6693 reticulata var subregularis 3319 DC Uitvlug
6694 arachnoidea var arachnoidea 3319 DA Kanetvlei t/o
6696 arachnoidea var arachnoidea 3319 BD W Osplaas tunnel
6697 arachnoidea var arachnoidea 3319 BD E Osplaas tunnel
6698 venosa ssp granulata 3319 BA Karoopoort
6700 arachnoidea var arachnoidea 3320 CA Soutkuil
6702 pulchella var pulchella 3320 CA Soutkuil
6703 arachnoidea var arachnoidea 3320 DA Bellair Dam
6704 arachnoidea/mucronata 3320 CB Ouberg, Below proteas
6705 arachnoidea/mucronata 3320 CB Ouberg, with E Enopla
6706 arachnoidea/mucronata 3320 CB Ouberg, with Pteronia paniculata]
6710 mirabilis var depauperata 3420 BB Boesmansrivier
6711 mirabilis var depauperata 3420 BB Boesmansrivier
6712 mirabilis var depauperata 3420 BB Boesmansrivier
6713 mirabilis var depauperata 3420 BB Boesmansrivier
6714 mirabilis var depauperata 3319 DC Koeniesriver
6719 pumila 3319 DD Dassiehoek, N Robertson
6720 mirabilis var depauperata 3319 DC Rietvleikloof
6721 reticulata var reticulata 3319 DD 12km W Robertson
6722 pumila 3319 DC Boschfontein
6723 mirabilis var depauperata 3319 DC Takkap
6724 emelyae var emelyae 3322 CD Klipdrif, 30km S Oudtshoorn
6725 pungens 3323 DD W Braamrivier
6727 outeniquensis 3322 CD Herold
6727 scabra var scabra 3323 DB Braamrivier
6728 scabra var scabra 3323 CC W DeVlugt, ALfreds Pass
6729 transiens 3323 CC just N De Vlugt
6730 mucronata var habdomadis 3321 Seweweekspoort
6732 scabra var starkiana 3322 AD Lentelus, Schoemanspoort
6737 reticulata var reticulata 3319 DC N Mowers Stn
6738 arachnoidea var arachnoidea 3319 DC N Mowers Stn
6739 arachnoidea var nigricans 3321 CC W Springfontein
6740 arachnoidea var aranea 3320 DD Kleindoringrivier
6741 emelyae var major 3320 DD Kleindoringrivier
6743 mucronata var mucronata 3320 DD 5km SSE Barrydale
6744 arachnoidea var setata 3320 DC Tradouw Pass
6745 floribunda var floribunda 3421 AB RoseInnis Drive
6746 chloracantha var denticulifera 3321 BB NWHerbertsdale
6747 magnifica var acuminata 3421 BD Vleesbaai
6749 turgida var suberecta 3421 BB Die Hoek, Gouritz Gorge
6750 minima var minima 3421 BD Melkhoutfontein
6751 magnifica var splendens 3421 AB Soutpan, W Albertinia
6756 herbacea var paynei 3319 DD S Olifantsdoorn
6757 mirabilis var depauperata 3419 BB Hoeksrivierkloof
6758 nortieri var nortieri 3219 CD 5km SE Sneeukop
6759 nortieri var nortieri 3219 CD 10km SE Sneeukop
6760 maraisii var maraisii 3319 DD Houtbaaiskloof
6761 maraisii var meiringii 3319 DD Top Bakenskop
6762 arachnoidea var setata 3320 DB Kuitfontein, NE Plathuis
6763 pulchella var globifera 3320 DB Wolwefontein, Plathuis
6764 blackburniae var blackburniae 3320 DB Wolwefontein, Plathuis
6767 outeniquensis 3322 CC Herold
6768 fasciata 3324 CD Moordenaarskloof
6769 viscosa 3324 CD Moordenaarskloof
6770 viscosaXfasciata 3324 CD Moordenaarskloof
6771 transiens 3324 CD Moordenaarskloof, at River
6772 pungens 3324 CC Joubertskraal
6773 cooperi var isabellae 3324 CC Joubertskraal
6774 scabra var scabra 3323 DD Braamrivier
6775 scabraXpungens 3323 DD Braamrivier
6776 cooperi var cooperi 3225 DA  SW Glen Avon
6777 nigra var nigra 3225 DA N Slagtersnek
6778 cooperi var pilifera 3225 DD New Slagtersnek
6780 pungens 3323 DD W Braamrivier
6781 maraisii var notabilis 3319 DD Bergplaas
6783 mirabilis var mirabilis 3419 DB Mierkraal
6784 cooperi var gordoniana 3324 DC Draaihoek, 10,5km SW Patensie
6786 cooperi var gordoniana 3324 DC Draaihoek, 10km SW Patensie
6788 viscosa 3324 DA Komdomo, Baviaanskl.
6789 cooperi var picturata 3324 DA 2km N Komdomo
6790 cooperi var picturata 3324 DC W Kranz, E Wistaria
6791 cooperi var picturata 3324 DC E Kranz, E Wistaria
6792 cooperi var gordoniana 3424 CB N Jeffrey’s Bay
6793 cooperi var gordoniana 3324 DD N Zuurbron
6794 fasciata 3324 DD Hankey Pass, N Zuurbron
6795 fasciata 3424 BB Kabeljouws
6796 fasciata 3424 BB N Jeffrey’s Bay
6797 fasciata 3424 BB W Bank Zeekoevlei River
6798 cooperi var isabellae 3324 DB Houtkloof, Elandsriver
6799 cooperi var gordoniana 3324 DD E Roodewal, W Patensie
6801 cooperi var isabellae 3324 DD Milton, SW Hankey
6802 cooperi var isabellae 3324 DD Philips Tunnel, Hankey
6803 attenuata var radula 3324 DD Hallelujah, N Hankey
6804 cooperi var gordoniana 3324 DD immed. NE Hankey
6805 cooperi var gordoniana 3324 DD N Hankey, Hallelujah
6806 fasciata 3325 CC Loerie Dam
6807 fasciata 3324 DD 2km E Hankey
6808 cooperi var isabellae 3325 CC E Gamtoos Bridge
6809 fasciata 3325 CC S Gamtoos Bridge
6810 cooperi var gordoniana 3323 DD N Joubertina
6811 cooperi var gordoniana 3323 CA Uniondale Poort
6813 pygmaea var pygmaea 3422 AA Moss Gas, W Mossel Bay
6815 maculata var maculata 3319 CB Audensberg Peak
6816 sordida var sordida 3325 BC SE. Kirkwood
6817 magnifica var atrofusca 3421 AA NW Kweekkraal farmhouse
6820 monticola var monticola 3322 DB W Uniondale, NE Kykoe to.
6821 scabra var scabra 3322 DB NE Kykoe to.
6822 viscosa 3323 DA Bottom Nuwekloof
6823 monticola var bavensis 3324 CA Top Bosrug
6824 cooperi var isabellae 3324 CA Koutnek Kougaberg
6825 transiens 3324 CA S Geelhoutboskloof camp
6826 cooperi var tenera 3324 CA Rooikloof
6827 cooperi var picturata 3324 CB E. Rooihoek campsite
6828 attenuata 3324 DA Sandlands Cambria
6830 cooperi var gordoniana 3324 DA E Komdomo
6831 attenuata 3324 DD Hallelujah, N Hankey
6832 cooperi var tenera 3326 BA E foot Plutosvale
6833 cooperi var tenera 3326 BA ft Plutosvale
6834 cooperi var tenera 3326 BA W ft Plutosvale
6835 cooperi var tenera 3326 BA WW ft Plutosvale
6836 cymbiformis var incurvula 3326 BA W Rolfs Rock, Plutosvale
6837 cymbiformis var incurvula 3326 BA S Rolfs Rock, Plutosvale
6838 cymbiformis var incurvula 3326 BA SE Rolfs Rock, Plutosvale
6839 cymbiformis var incurvula 3326 BA W Mindmill, Plutosvale
6840 cooperi var tenera 3326 BA opposite windmill, mid Plutosvale
6841 attenuata 3326 BA Rolfs Rock Plutosvale
6842 bolusii var blackbeardiana 3227 BC Inverbolo
6843 cymbiformis var reddii 3227 BC Inverbolo
6844 angustifolia 3326 AC E Alicedale
6845 cooperi var cooperi 3326 AC E Alicedale
6847 cymbiformis var obtusa 3326 AC SW Alicedale
6848 cymbiformis var obtusa 3326 AC NW Alicedale
6849 angustifolia 3325 BB Aalwynspoort Stn
6850 cymbiformis var obtusa 3325 BB Swartwaterpoort
6851 aristata 3325 BB Modderfontein, S Verdun
6852 aristata 3325 BA Stonefountain
6855 decipiens var virella 3325 AA E Waterford Bridge
6856 arachnoidea var xiphiophylla 3323 BB NE Fullarton Siding
6858 glauca var herrei 3323 BB NE Fullarton Siding
6859 floribunda var floribunda 3420 AB SW Swellendam
6860 maraisii/mirabilis 3420 AB 4km SW Swellendam
6861 maraisii/heidelbergenis 3420 AB SW Swellendam, Bredasdorp to.
6862 mirabilis/maraisii 3420 AA NW Rondeheuwel, Stormsvlei
6863 mirabilis var diversicolor 3419 BB Boesmansriv/Olifantsdoorn
6864 mirabilis var depauperata 3319 DD Whipstock, W McGregor
6865 mirabilis var consanguinea 3319 DD Dwarswaterkloof, W McGregor
6867 arachnoidea var arachnoidea 3320 CB 5km NE Ouberg farmhouse
6868 arachnoidea var arachnoidea 3320 CD W Twistniet, E Montagu
6871 arachnoidea var arachnoidea 3319 DD Pietersfontein, NW Montagu
6873 arachnoidea var arachnoidea 3319 DD NW Pietersfontein, Montagu
6875 maraisii var maraisii 3320 CC NE Ashton. Cogmanskloof
6876 mirabilis var depauperata 3419 BB E Olifantsdoornkloof
6877 maraisii var maraisii 3319 DD Die Nekkie, W Robertson
6878 maraisii/mirabilis 3419 BB SW Swellendam
6879 maraisii/floribunda 3420 BA Koppies, SE Swellendam
6880 rossouwii 3420 BA Oudekraalkop
6881 floribunda 3420 BA S Oudekraalkop
6882 heidlbergensis 3420 AD W Juliesfontein
6883 mucronata var mucronata 3320 CD Poortjieskloof Dam
6884 cymbiformis var incurvula 3326 BA Rolf’s Rock, N Plutosvale
6885 decipiens var cyanea 3222 DC Trakaskuilen, S Beaufort West]
6886 heidelbergensis/mirabilis 3420 AD N Juliesfnt, S Brakfnt.
6888 variegata var modesta 3420 BC NW Slopes Potberg
6889 heidelbergensis var scabra 3420 BC NW slopes Potberg
6890 heidelbergensis var scabra 3420 AD Witkop, N Brakfontein
6891 cooperi var tenera 3326 BA 300m W lower Roff’s Rock windmill
6892 cymbiformis var incurvula 3326 BA below 6891, Roff’s Rock
6893 cymbiformis var incurvula 3326 BA 400m W Roff’s Rock
6894 angustifolia 3326 AB Thornkloof, NW Grahamstown
6895 cymbiformis var obtusa 3326 AB SE Thornkloof house
6896 coarctata var adelaidensis 3326 AB SE Thornkloof house
6897 aristata 3325 BB SE Commadagga house
6899 aristata 3325 AB Paddafontein crossing
6901 aristata 3325 AB 400m W Hopewell farm
6903 cymbiformis var cymbiformis 3325 AD S slopes Spekboomkop
6904 cooperi var gordoniana 3325 AD Top mnt. Wilgerfontein
6905 cooperi var isabellae 3325 AD Top mnt. Wilgerfontein
6907 glauca/coarctata 3325 AD Top mnt. Wilgerfontein
6908 glauca 3325 AD SE DePlaat
6909 cooperi var gracilis 3325 AD SE DePlaat
6910 cooperi var isabellae 3325 AD SE DePlaat
6911 cooperi var gracilis 3325 AD Kaboegapoort, Cliff face
6914 cooperi var gracilis 3325 AD East Spekboomberg
6915 cooperi var gracilis 3325 AD SE above, Koch’s Dam
6916 aristata 3325 AB Spitzkop, Kaboega
6917 aristata 3325 AB Original Kaboega gate
6920 aristata 3325 DA Soutkloof
6921 cooperi var doldii 3327 BA Chalumna crossing
6922 cooperi var pilifera 3325 BA Oudekraal
6924 cooperi var gracilis 3325 AD Kaboega Dam
6925 cymbiformis var cymbiformis 3325 AD Kok se Pad
6927 bolusii var pringlei 3325 BB 3km W Ripon Stn
6928 cooperi var isabellae 3324 DC Nuwelande, Patensie
6929 longiana 3324 DC Nuwelande, Patensie
6930 cooperi var picturata 3324 DC E. Andrieskraal
6932 cooperi var isabellae 3424 BB Krom River, Ripon
6933 cooperi var gordoniana 3424 BB 7km E Humansdorp
6934 fasciata 3424 BB 4km SW Humansdorp
6935 cooperi var gracilis 3325 AD S DePlaat
6936 cooperi var gordoniana 3323 CA N Uniondale
6937 decipiens var scottiana 3323 CA Vetvlei, N Uniondale
6938 decipiens var scottiana 3323 CA Woedjieskloof, NE Uniondale
6939 decipiens var scottiana 3323 AD Ghwarriepoort, Keurfontein
6940 cooperi var gracilis 3325 AB Klipfontein, E DePlaat
6942 aristata 3325 AB DePlaat Weir
6943 cooperi var gracilis 3325 AD Dassiekop
6945 nigra 3325 AB N DePlaat house
6947 sordida 3325 AD S DePlaat house
6949 cooperi var pilifera 3325 AD Vyeboomfontein
6951 maraisii var 3320 CC NN Ashton
6952 marginata 3320 CC N Ashton
6953 maraisii var maraisii 3319 DD Le Chasseur Nek, S Sandberg
6954 maraisii var maraisii 3420 AA N Stormsvlei
6955 mirabilis var triebneriana 3420 AA Stormsvlei Pass
6956 maraisii var maraisii 3319 DD SWslopes,Rooiberg, Eilandia
6957 reticulata 3319 DD S valley Rooiberg
6958 pumila 3319 DD S slopes Rooiberg
6959 maraisii var maraisii 3319 DD Upper SE slopes, Rooiberg
6959 maraisii var maraisii 3319 DD Kleinspitzkop, Vrolikheid
6960 maraisii var maraisii 3320 CC W Sanddrift
6961 maraisii var maraisii 3319 DD Muiskraalkop, Robertson
6962 maraisii var maraisii 3320 CC Goudmyn, E river
6963 maraisii var maraisii 3320 CC 5km W Bonnievale
6964 maraisii var maraisii 3320 CC 5km E Zandvliet
6965 maraisii var maraisii 3320 CC 2km E Zandvliet
6966 maraisii var maraisii 3319 DD Steenbokshoogte, Vrolikheid
6967 maraisii var maraisii 3319 DD W Kanonkop, Vrolikheid
6968 maraisii var maraisii 3319 DD Witkranz, Vrolikheid
6969 maraisii var maraisii 3319 DD Kleinspitzkop, Vrolikheid
6970 herbacea var flaccida 3319 DD Rooiberg, Vinkrivier
6971 heidelbergensis/maraisii 3320 CC Jakkalshoek, E Bonnievale
6972 maraisii 3319 DD with flaccida, Rooiberg
6973 maraisii var maraisii 3419 BD N Napier
6974 heidelbergensis var scabra 3320 CC Tonteldooskop
6976 heidelbergensis var scabra 3320 CC S Tonteldooskop
6977 heidelbergensis var scabra 3320 CC south aspect at 6974
6978 heidelbergensis var scabra 3320 CC between 6974/6977
6979 heidelbergensis/maraisii 3320 CC Loerkop, E. Ashton
6981 heidelbergensis/maraisii 3320 CC SE Loerkop
6982 mutica var mutica 3420 AC Hasiesdrift
6983 rossouwii var rossouwii 3420 CA Soutkloof, NW Bredsadorp
6984 rossouwii var rossouwii 3420 CA Nooitgedacht, NW Bredasdorp
6985 rossouwii var calcarea 3420 AD DeHoop at cottages
6986 rossouwii var petrophila 3420 CA Karsriver W Limeworks
6987 mirabilis var badia 3419 BD Sandfontein, W Napier
6988 pumila 3319 DD Pietersfontein, NW Montagu
6988 heidelbergensis var scabra 3320 CC Nooitgedacht at Farmhouse
6989 heidelbergensis var scabra 3320 CC 2km SE Nooitgedacht farmhouse
6990 heidelbergensis var scabra 3320 CC 2km NE Mardouw farmhse.
6991 mucronata 3320 DA Jakkalsfontein
6992 heidelbergensis var scabra 3320 CD Langverwacht
6994 arachnoidea var nigricans 3321 DB NE Tierkloof, Gamka
6995 viscosa 3321 DB E. Tierkloof
6996 arachnoidea var setata 3321 DB SE Tierkloof
6997 viscosa 3321 DB S Tierkloof
6998 arachnoidea var setata 3321 DA Top Rooiberg
7001 arachnoidea var setata 3321 DA Groenefontein
7002 viscosa 3321 DA Groenefontein
7003 arachnoidea var setata 3321 DA W Groenefontein
7006 bolusii var pringlei 3225 CC Moederzoonpoort
7008 bolusii var pringlei 3225 CC Rietvlei NE Waterford
7010 aristata 3325 AB SE DePlaat Weir
7011 aristata 3325 AB SE DePlaat Weir
7012 aristata 3325 AB Buffelsnek, Kaboega
7014 truncata 3322 CB N Dysseldorp
7017 cooperi var isabellae 3325 BC S Klipfontein
7021 bolusii var bolusii 3225 CC SE Pearston
7022 decipiens var virella 3225 CC SE Pearston
7023 decipiens var virella 3224 DB SW Pearston
7024 viscosa 3322 AB Sleutelfontein, Prince Albert
7025 decipiens var cyanea 3222 DC Trakaskuilen
7026 decipiens var decipiens 3322 BC Klaarstroom
7028 decipiens var virella 3325 AA Darlington Dam
7029 mutica var mutica 3420 AD 25km SW Swellendam
7030 maraisii var maraisii 3420 AB 22km SW Swellendam
7031 mutica var mutica 3420 ab 18km SW Swellendam
7032 mutica var mutica 3420 AA Rietfontein, S Stormsvlei
7033 arachnoidea 3319 DD W. Cape Lime works, Mowers
7035 kingiana/minima 3321 AD Rooiberg, S Calitzdorp
7036 heidelbergensis var scabra 3320 CC Witkop, SW Janharmsgat
7037 heidelbergensis var scabra 3320 CC Middelrivier, Drew
7038 marginataXpumila 3320 CC NW Sanddrift
7041 decipiens var virella 3224 DD Palmietfontein, NE Jansenville
7042 bolusii var bolusii 3224 DD Hinchenbrook
7043 decipiens var virella 3224 DD Langollen
7046 bolusii var bolusii 3224 DD DeRust, S Lootskloof.
7047 decipiens var virella 3224 DD DeRust, S Lootskloof
7049 cooperi var pilifera 3325 AD SE Klipfontein, Kaboega
7051 cooperi var gracilis 3326 AB Helspoort, upper W facing
7052 cooperi var gracilis 3326 AB Helspoort, lower east
7053 cooperi var gracilis 3326 AB Helspoort, lower west
7054 cooperi var gracilis 3326 AB Helspoort, upper east
7055 maraisii var nov 3420 CC Rietvlei (Shielam)
7056 heidelbergensis va scabr 3420 CC NW Sandrift, Drew
7057 heidelbergensis var scabra 3420 CC W Sanddrift, Drew
7058 pumilaXmarginata 3420 CC S Sanddrift, Drew
7059 mirabilis var triebneriana 3419 BC Jongensklip
7060 mutica var mutica 3420 AD Beyersdal
7061 variegata var modesta 3420 AD Kathoek
7063 heidelbergensis 3420 AD SE Beyersdal
7064 mutica var mutica 3320 CC Sanddrift
7065 pubescens var pubescens 3319 DC Sandberg
7066 pubescens var livida 3319 CD Lemoenpoort
7067 herbacea 3319 DC Sandberg
7068 arachnoidea var scabrispina 3320 BB Witteberg Rd
7069 viscosa 3320 BB Witteberg Rd
7070 nortieri var pehlemanniae 3320 BB NW Laingsburg
7071 arachnoidea var scabrispina 3320 BB NW Laingsburg
7072 marumiana var dimorpha Konstabel
7073 wittebergensis S Laingsburg
7074 mirabilis var triebneriana 3419 BB Verdwaalskloof, Riviersonderend
7075 mutica var mutica 3420 DD Grootvlakte, E Riviersonderend
7076 herbacea var paynei 3419 DD S Grootrivier
7077 maraisii var maraisii 3319 DD Wolwedrif, SW Bonnievale
7078 pumila 3319 DD Wolwedrif, SW Bonnievale
7079 maraisii var meiringii 3319 DD W Middelplaas
7080 maraisii var meiringii 3320 CC W Wakkerstroom
7081 maraisii var meiringii 3320 CC E Goudmyn
7082 maraisii var meiringii 3320 CC Bobbejaanskloof
7083 maraisii var meiringii 3320 CC NE Goudmyn
7084 maraisii var meiringii 3320 CC E Olive grove
7085 maraisii var meiringii 3320 CC Station Hill
7086 maraisii var maraisii 3420 AA SE Bokdam
7087 mirabilis var triebneriana 3420 AA Brakfontein
7088 reticulata var reticulata 3319 DD W Middelplaas
7089 pumila 3320 CC E Bonnievale
7090 mirabilis var mirabilis 3419 DB Mierkraal
7091 mirabilis var triebneriana 3419 BD NW Napier
7092 mirabilis var triebneriana 3420 AB Elandskloof
7096 pumila 3319 DD Koningsriver
7097 maraisii var notabilis 3319 DD Kranzkop
7098 maraisii var notabilis 3319 DD Bergplaas
7099 heidelbergensis 3420 AA Nuutbegin
7100 minima 3420 AA Nuutbegin
7101 variegata 3321 AB Hoekraal
7102 magnifica var splendens 3321 BA Welgevonden
7103 floribunda var dentata 3321 BA N Ouvloer
7104 floribunda var dentata 3321 BA NE Ouvloer
7105 turgida var suberecta 3321 BA NE Ouvloer
7106 floribunda var dentata 3321 AB Snymanskraal
7107 retusa 3321 AA NE Melkboom
7108 heidelbergensis 3321 AA NE Melkboom
7109 heidelbergensis 3321 AA Klien Kragga
7110 retusa 3321 AA N Melkboom
7111 heidelbergensis 3321 AA N Melkboom
7112 heidelbergensis 3321 AA NW Kweekkraal
7113 heidelbergensis 3321 AA NW Kweekkraal
7114 heidelbergensis 3321 AA NW Kweekkraal
7115 heidelbergensis 3321 AA NW Kweekkraal
7116 heidelbergensis 3321 AA NW Kweekkraal
7117 heidelbergensis 3321 AA NW Kweekkraal
7118 heidelbergensis 3321 AA NW Kweekkraal
7121 cymbiformis 3325 BD Gatjewolf
7125 attenuata 3325 BC N Enon
7126 cooperi var puberula 3325 AD Wilgerfontein
7127 glauca 3325 AD Buffelsnek
7128 angustifolia var baylissii 3325 BD Wells Gate
7130 venosa subsp granulata 3319 BA Karoopoort
7131 venosa subsp granulata 3220 CC E Bizansgat
7133 mutica var nigra 3420 BB Goedehoop
7134 mutica var nigra 3420 BB Klipdrift
7135 venosa subsp venosa 3420 BB Sandkraal
7136 limifolia Bridgewater Nursery
7137 limifolia 2732 CD Pumalanga
7138 limifolia 2732 CB Inxwala, Mkhuze
7139 limifolia 2731 BC Ncotshane, Pongola
7140 limifolia 2731 BC Draaiwater, Pakamisa
7141 limifolia 2531 CC N Rymers Creek
7142 limifolia 2531 CC Mays Mine
7143 limifolia 2531 CB Boondoks
7144 limifolia 2531 CB N Manders Dam
7145 limifolia 2531 CB Three Sisters
7146 limifolia 2731 CA Bivaan Dam
7147 limifolia 2731 CB Ithala
7148 limifolia 2731 AD Ithala
7149 viscosa N.Koup Stn
7150 decipiens var cyanea W.Merweville
7151 heidelbergensis 3421 AA N Kweekkraal W
7152 heidelbergensis 3421 AA N Kweekkraal W
7153 heidelbergensis 3421 AA N Kweekkraal W
7154 heidelbergensis 3421 AA N Kweekkraal W
7155 floribunda 3421 AA SW Pretoriuskop
7156 floribunda 3421 AA W Pretoriuskop
7157 atrofusca 3421 AA N Bosgasiekop
7158 floribunda 3421 AA N Bosgasiekop
7159 atrofusca/floribunda 3421 AA N Bosgasiekop
7160 magnifica 3421 AA N Kweekkraal E
7161 magnifica 3421 AA N Kweekkraal E
7162 magnifica 3421 AA N Kweekkraal E
7163 magnifica 3421 AA S Riversdale t.loc
7164 floribunda cf. 3420 BD Dassieklip
7165 heidelbergensis 3420 BB Matjestoon
7166 pumila 3319 BD Mowers Stn
7167 pumila 3319 BC Apiesklip
7168 cooperi var 3325 AD Kabouga
7169 attenuata 3325 BC Enon
7170 cymbiformis var 3325 BC Enon
7171 glauca/coarctata 3325 AD S Wilgefontein
7172 floribunda 3421 AA N Kweekkraal E
7173 retusa 3421 AA N Kweekkraal E
7174 maraisii 3319 DD Skurweberg
7175 pumila 3319 DD W Highlands, Goree
7176 maraisii 3319 DD N Agtervink
7178 viscosa 3323 BD Ruiterspoort
7179 viscosa 3323 BD Constantia
7180 decipiens 3323 BD Constantia
7182 monticola 3323 BC SW Koegoeroe
7183 momticola 3323 BC SE Koegoeroe
7184 bayeri 3323 CA Uniondale Fort
7185 monticola 3323 CA Uniondale Height
7188 decipiens/scottiana 3323 CA 5km E Hoekplaas
7189 decipiens/scottiana 3323 CA 2km NW Hoekplaas
7190 mucronata 3322 BD 3km E Rooiloop Stn.
7193 cf scottiana 3322 BC 5km W Rooiloop Stn
7194 arachnoidea 3322 DA E Middelplaas Stn
7195 arachnoidea 3322 BC 2km E Vlakteplaas
7196 bayeri 3322 BC SW Derust
7200 truncata 3322 CA NE Oudtshoorn
7201 truncata 3322 CA Volmoed
7202 morrisiae 3322 CA Volmoed
7203 arachnoidea 3321 CB Ladismith
7204 scabra 3322 DA SE Doornkloof
7205 arachnoidea var nigricans 3321 DA W Vensterkranz.
7206 arachnoidea var nigricans 3321 CA Buffelsdrif S
7207 arachnoidea var nigricans 3321 CA Buffelsdrif
7208 arachnoidea var nigricans 3321 CA Bakenskop
7210 arachnoidea var nigricans 3321 CA E Vensterkrans
7211 arachnoidea var nigricans 3321 CC W Springfontein
7212 mucronata var mucronata 3321 CC Tradouwshoek
7213 mucronata var mucronata 3321 CC S Barrydale S aspect
7214 maraisii 3321 CC S Barrydale S aspect
7216 mucronata var mucronata 3321 CC S Barrydale
7217 heidelbergensis/magnifica 3421 AA Rooikop Melkboom
7218 floribunda/magnifica 3420 BB Duiwenhoks Morn Star
7219 turgida 3420 BB Duiwenhoks Morn Star
7220 floribunda/magnifica 3420 BB NE Morning Star
7221 mutica var nigra 3420 BB NE Morning Star
7223 scabra 3322 DA Diepkloof, E De Rust
7224 floribunda/chloracantha 3421 BB Cooper Siding
7225 retusa 3421 AA S Droekloof Riversd.
7226 turgida 3420 BB Witheuwel Pienaarsr
7227 heidelbergensis/magnifica 3420 BB Witheuwel Pienaarsr
7228 turgida 3420 BB Somona S Heidelberg
7229 heidelbergensis/magnifica 3420 BB Somona S Heidelberg
7231 bayer1 (seed) 3322 DA Kleingeluk, E DeRust
7232 turgida/heidelbergenss 3420 BB S Heidelberg Reserve
7233 turgida/heidelbergenss 3420 BB Hooikraal (Die Plotte N)Bayer
7234 turgida/heidelbergenss 3420 BB Hooikraal (Die Plotte S)
7236 turgida/retusa 3420 BB E Heidelberg Dam
7237 magnifica cf.var atrofusca 3420 BB Andrieskraal/Skeideing
7238 magnifica cf.var atrofusca 3420 BB Andrieskraal/Skeiding
7239 magnifica cf.var atrofusca 3420 BB Andrieskraal/Skeiding
7240 turgida/retusa 3420 BB Skeiding
7241 maraisii 3419 DD Voordieberg, McGregor
7242 mirabilis/maraisii 3420 AA Witkop, N Uitvlug Annex
7243 mirabilis/maraisii 3420 AA SE Uitvlug Annex
7244 mirabilis/maraisii 3420 AB E Die Kop, SE sh345
7245 mirabilis/maraisii 3420 AB do E sh345
7246 mutica 3420 AC Verbrandskloof
7247 mirabilis/maraisii 3419 BD 0.5km NE Napier
7248 mirabilis/maraisii 3420 DB Ballyfar N, Infanta
7249 mirabilis/maraisii 3420 DB Ballyfar S, Infanta
7250 maraisii/heidelbergensis 3420 BC Uitrus (Whisky Creek)
7251 mirabilis 3420 BC Sandhoogte, DeHoop
7252 mirabilis 3420 AA NE Klipfontein
7253 mirabilis 3420 AA N Klipfontein
7254 mirabilis 3420 AA NW Noukloof Homestead
7255 elizeae 3420 AA 28km E Riviersonderend
7257 maraisii 3420 AA SW Stormsvlei homestead
7258 minima 3420 BC Uitrus (Whisky Creek)
7259 mirabilis 3419 BD Napier Stn
7260 mirabilis 3419 BD S Mierkraal, Napier
7261 mirabilis 3420 AC NW Bredasdorp
7262 mirabilis 3420 BA Ouplaas, SE Greyton
7263 maraisii 3319 DC Droerivierberg B
7264 maraisii 3319 DC Droerivierberg A
7265 maraisii 3319 DC Haumanskloof
7266 pubescens var livida 3319 CD E Lemoenpoort S
7268 herbacea 3319 CD E Lemoenpoort S
7269 maraisii v meiringii 3320 CC Edendale, Bonnievale
7270 maculata 3319 CD Ouhoek Mt Moddergat
7271 maculata 3319 CB S Kwaggaskloof Dam
7273 herbacea 3319 CD Draaivlei Kop
7280 mirabilis 3419 BD N Mierkraal Napier
7283 mirabiis var 3420 AC W Adoonskop
7285 mirabiis var 3420 AC N Brakkloof
7287 mirabiis var 3420 AC SW Adoonskop roadside
7288 mirabilis var 3419 BC N Diepkloof, Oudebakho.
7295 mirabilis var 3419 BC N Diepkloof, Oudebakho.
7299 rossouwii 3420 AC NW Soutkloof
7325 maraisii 3320 CC Laastewater
7327 maraisii 3320 CC 4km E Bonnievale
7334 maraisii 3420 CA 4km NE Bredasdorp
7337 heidelbergensis var scabra 3420 BC SE Brakfontein
7356 mirabilis var pilosa 3420 BC NE Buffelsfontein
7374 decipiens cf. var. minor 3322 AC Malvadraai, Swartberg.
7375 decipiens var. decipiens 3322 AC Scholtzkloof.
7376 marumiana cf var. viridis 3322 AC Scholtzkloof.
7377 viscosa 3322 AC Scholtzkloof.
7382 nortieri var. devriesii 3322 AC N Prince Albert
7384 viscosa 3322 AC Syferfontein
7386 nigra 3322 AC Toekomsrus
7417 floribunda/chlorcantha 3421 BD Melkhoutfontein
7419 venosa var. venosa 3421 BA Die Eiland Albertinia
7420 chloracantha var.choracantha 3421 BA Die Eiland Albertinia
7425 chloracantha var.choracantha 3422 AA Wolwedans Dam
7453 marginataXminima 3420 CA NE Bredasdorp
7485 herbacea 3319 DC Droogeriviersberg
7486 cooperi var.pilifera 3325 AB Vygeboomfontein
7487 floribunda var. floribunda 3420 BC Byeneskop
7488 minima 3420 BC Byeneskop
7492 floribunda var. floribunda 3420 BC SE Klipfontein Potberg
7494 floribunda var. floribunda 3420 BC SE above
7495 floribunda var. scabra? 3420 BC SE above
7497 maraisii cf. atrofusca 3420 BC SE Kleinberg
7499 floribunda/heidelbergensis 3420 BC SW Byeneskop
7500 mirabilis/mutica 3420 AD Die Kop E
7502 mirabilis/maraisii 3420 AD Die Kop W
7503 mrabilis/maraisii 3420 AD Die Kop mid
7504 mirabilis/heidelbergensis 3420 AD Die Kop N
7507 magnifica var fusca 3421 AD W Albertinia
7508 magnifica var. dekenahii 3421 BA Draaihoek
7509 floribunda var dentata 3421 BA Draaihoek
7510 maraisii var. maraisii 3420 CA NE Bredasdorp
7511 variegata var. variegata 3421 BA Swartklip
7512 turgida var. suberecta 3421 BA Draaihoek
7513 maraisii/heidelbergensis 3420 AA Klipfontein Vaan+F1269drigsdrift
7514 mirabilis 3420 AD Stoffelsriver
7515 variegata 3420 AD Stoffelsriver
7516 mirabilis/floribunda 3420 AD W Kleinberg, Malgas
7518 variegata 3420 AD S Kleinberg
7519 minima 3420 AD S Kleinberg
7520 mirabilis 3420 AD NW Stoffelsriver
7522 nortieri 3118 BC Arizona
7523 nortieri 3118 DD Giftberg
7524 nortier 3118 BB N Clanwilliam
7525 nortieri 3118 BB S Clanwilliam
7526 maculata 3319 CD Nekkies W
7527 mucronata 3320 DD Orange Grove S Barrydale
7577 arachnoidea 3119 AD Nuweplaas NWDville
7578 nortieri 3119 AD Nuweplaas NWDville
7579 nortieri 3118 AC Trawal Bridge
7580 nortieri 3219 CB Bakkamerfontein
7586 cooperi gordoniana Kliprivier Uniondale
7593 aristata DePlaat to
7594 nortieri albispina Koup
7595 nortieri 3219 AC N Dwarsriver
7597 nortieri 3219 CA Kunje
7599 nortieri 3219 CA SE Kunje
7604 minima 3420 CA NW Bredasdorp
7605 mirabilis 3420 CA NW Bredasdorp
7608 mirabilis ‘pilosa’ 3420 BC Melkhoutrivier
7609 mirabilis ‘pilosa’ 3420 BC Melkhoutrivier
7612 mirabilis 3420 AB Diamant W
7615 mirabilis 3420 BC Aalwee Malgas
7622 mirabilis 3420 CC Bergendal Ashton
7623 mirabilis 3420 CC Sarahsriver
7634 cooperi/cymbiformis 3225 DA Glen Avon
7635 sordida Swartkop Kaboega
7636 cooperi/cymbiformis N Swartkop Kaboega
7640 granulata Oskopvlakte
7641 viscosa Blokhuis
7642 wittebergensis Ezelsfontein
7644 arachnoidea Bakoven
7646 arachnoidea Bizamsgat
7658 arachnoidea SW Perdekraal
7659 ganulata Bantamsfontein
7660 arachnoidea Bantamsfontein NW
7666 arachnoidea Patatsriver
7670 arachnoidea Keurkloof
7671 arachnoidea Bulhouer
7672 arachnoidea NW Bulhouer
7673 arachnoidea N Bulhouer
7676 arachnoidea 8km N Bulhouer
7677 arachnoidea Dwarsindieweg
7678 arachnoidea SW Josefskraal
7679 arachnoidea N Josefskraal
7681 marumiana Lospersberg
7686 arachnoidea N Dwarsindieweg
7688 arachnoidea Klein Tafelkop
7692 nortieri? Volstruisfontein
7693 arachnoidea Volstruisfontein
7695 glauca 3325 AD SW Darlington Dam
7697 aristata 3325 AD 2km N Swartkops
7698 aristata 3325 AD E Buffelsnek
7701 cooperi ‘puberula’ 3325 AD S Klipfontein
7702 glauca 3325 AD S Klipfontein
7703 aristata 7697
7704 mirabilis 3420 AB Bontebok Pk SE Office
7705 mirabilis’atrofusca’ 3420 BA Uitvlugt NW
7706 mirabilis ‘atrofusca’ 3420 BA Uitvlugt NNW
7707 retusa ‘nigra’ 3420 BA Goedverwagting
7708 floribunda 3420 BA Goedverwagting
7709 rossouwii 3420 BA SW Grootkloof
7710 rossouwii 3420 BA WSW Grootkloof
7711 rossouwii 3420 BA NE Grootkloof
7713 mirabilis ‘atrofusca’ 3420 BA W Uitvlugt
7714 mirabilis ‘atrofusca’ 3420 BA E Uitvlugt
7715 mirabilis ‘atrofusca’ 3420 BA N Uitvlugt
7716 rossouwii 3420 BA E Uitvlugt
7717 mirabilis 3420 BA SW Koppies
7719 mirabilis 3420 AB S Dagbreek
7722 floribunda 3420 AB Appelbos
7723 mirabilis 3420 BA S Koppies
7724 mirabilis ‘paradoxa’ 3421 AC NE Oshoek
7726 mirabilis ‘paradoxa’ 3421 AC SE Vermaaklikheid
7727 mirabilis ‘paradoxa’ 3421 AC Koenserus
7728 mirabilis 3420 BB NE Lilliendal
7729 mirabilis 3420 BB NW Lilliendal
7732 rossouwii 3420 BC NW Stuurmanskraal
7733 rossouwii 3420 BC N Stuurmanskraal
7734 mirabilis 3420 BC Stuurmanskraal
7735 marginata 3420 BA E Koppies
7737 minima 3420 BA W Koppies
7738 floribunda major 3420 AB S Swellendam
7739 herbacea 3319 CB Glen Heatlie
7740 pumila 3319 CB Glen Heatlie
7741 mutica 3420 AB Dankbaar
7742 mirabilis 3420 AB Dankbaar
7743 minima 3420 AB Bontebok Park
7744 mirabilis 3420 AB Bontebok Park SW
7747 minima 3420 BD Sandfontein, N Witsand
7749 mirabilis 3420 BC Kadies Landing
7751 mirabilis 3420 BC Kadies Landing
7753 mirabilis 3420 BD Brakkuil
7754 retusa turgida 3420 BD Brakkuil
7756 retusa turgida 3420 BD Dassieklip
7757 minima 3421 AB Klipheuwel
7758 retusa 3421 AB Skietbaan Rdal.
7760 floribunda 3421 AB Witkleikop
7761 mirabilis 3421 AB Plattekop farm
7762 mirabilis 3421 AB W Plattekop
7763 mirabilis 3421 AB E Plattekop
7764 floribunda 3421 AB NW Platkop
7765 mirabilis 3421 AB NNW Plattekop
7765a retusa 3421 AB Plaatjieskop
7767 floribunda 3421 AB Platjieskop
7769 mirabilis 3421 AB Toringskop
7770 mirabilis 3421 AB W Soetmelksrivier
7771 retusa 3421 AB W Soetmelksrivier
7772 retusa 3421 AB Platkop Annexe
7774 floribunda ‘major’ 3420 AB S Swellendam
7776 retusa 3421 AA Pienaarsrivier
7778 mirabilis 3421 AB Komserante
7779 mirabilis 3421 AB Komserante
7780 retusa 3421 AB Komserante
7781 retusa 3421 AB Komserante
7782 minima 3421 AB Komserante
7783 mirabilis 3421 AB W Soetmelksrivier
7784 mirabilis 3421 AB W Soetmelksrivier
7785 retusa 3421 AB W Soetmelksrivier
7786 mirabilis 3421 AB W Soetmelksrivier
7790 retusa 3421 AB Groothoogtekop
7791 minima 3421 AB Groothoogtekop
7792 floribunda 3421 AB Groothoogtekop
7794 retusa nigra 3421 BB Droerivier
7797 limifolia 2632 CA E. Isiteki
7801 mutica 3420 BA Mullersrus
7803 rossouwii 3420 BB Morning Star
7804 retusa nigra 3420 BB Kransriviermond
7805 mirabilis 3420 AB Bontebok Park N
7807 minima 3420 BB Diepkloof
7808 mirabilis 3420 BB Diepkloof
7809 mirabilis 3420 BB Koeisekop
7810 retusa turgida 3420 BB Tierkloof
7811 mirabilis 3420 BB Kransriviermond
7812 retusa turgida 3420 BB Kransriviermond
7813 minima 3420 BB Skeiding
7814 mirabilis atrofusca 3420 BB Skeiding
7815 retusa turgida 3421 AB Klipdrift
7816 retusa turgida 3421 AB Heuningfontein
7817 variegata 3421 AB Klipfontein
7818 mirabilis 3421 AB Windsor
7819 marumiana dimorpha 3320 AD W Konstabel
7820 mirabilis jakubi 3421 AB Klipfontein
7821 rossouwi minor 3420 CA Rooivlei
7822 mirabilis 3420 CA Rooivlei
7823 mirabilis 3420 BA Klipbult
7825 chloracantha 3421 BB N Herbertsdale
7827 chloracantha 3421 BB S Herbertsdale
7828 parksiana 3422 AA Rooiheuwel
7829 chloracantha 3422 AA Rooiheuwel
7835 kingiana 3422 AA Rooiheuwel
7844 emelyae 3321 CC Brandrivier
7845 minima 3321 CC Brandrivier
7846 emelyae 3321 CC W Springfontein
7847 emelyae 3320 DD Kleindoringrivier
7848 emelyae 3321 CC E Springfontein
7849 retusa turgida 3321 DC Towerland
7850 emelyae 3321 DC Aasvoelvallei
7851 arachnoidea nigricans 3320 DD Kleindoornrivier
7856 emelyae major S Muiskraal
7857 emelyae multifolia W Muisktraal
7858 emelyae multifolia NW Muiskraal
7859 emelyae multifolia N Muiskraal.
7860 emelyae major 3km ENE Onverwacht
7861 emelyae W Zandkraal
7863 arachnoidea Orange Grove
7865 arachnoidea cf Keurkloof De Doorns
7866 chloracantha 3km E Herbertsdale
7867 chloracantha S above
7868 kingiana 3km E Herbertsdale
7869 monticola Williamsburg
7870 cooperi Williamsburg
7873 parksiana W Botlierskop
7876 mirabilis Dysselfontein SW Swellendam
7877 venosa var. venosa Doringrivier
7882 mirabilis ‘meiringii’ Edendale, Bonnievale
7886 mirabilis N Sandhoogte
7887 mirabilis W Rotterdam
7888 mutica SW Rotterdam
7889 mutica W Rotterdam
7890 mutica NW Rotterdfam
7891 minima W Rotterdam
7892 marginata W Rotterdam
7895 turgida Diepkloof, Malgas
7896 retusa nigra Heuningklip Tradouw
7897 retusa nigra Heuningklip Tradouw
7898 retusa nigra Heuningklip Tradouw
7899 mirabilis Heuningklip Tradouw
7901 mirabilis Felix Unite
7902 mirabilis Felix Unite
7903 mirabilis Felix Unite
7904 mirabilis Felix Unite
7905 mirabilis Felix Unite
7906 mirabilis Felix Unite
7907 mirabilis Felix Unite
7908 mirabilis Felix Unite
7910 floribunda Rietkuil
7912 mirabilis Rietkuil
7913 mirabilis Rietkuil
7914 mirabilis Van Reenens Crest
7915 mirabilis Van Reenens Crest
7916 mirabilis Van Reenens Crest
7917 mirabilis Van Reenens Crest
7918 mirabilis Van Reenens Crest
7919 mirabilis Van Reenens Crest
7920 retusa var nigra Van Reenens Crest
7921 retusa var nigra Van Reenens Crest
7922 mirabilis Diepkloof NW
7923 variegata Diepkloof
7924 variegata Diepkloof
7925 variegata Diepkloof.
7926 minima Diepkloof
7927 variegata Diepkloof
7928 minima Diepkloof
7930 minima W Diepkloof
7931 mirabilis S 7516 Kleinberg
7932 minima S Kleinberg
7933 mirabilis S7496 Se Kleinberg
7934 mutica Wolwefontein
7935 mutica W Crodini
7936 mirabilis Langvlei
7937 mutica Platkop
7938 mirabilis SE Stoffelsriver
7941 mutica Ouplaas, SE Greyton
7943 mutica E. Klipbankskloof
7944 mirabilis SE Klipbankskloof
7945 mutica SE Klipbankskloof
7949 marginata Middelrivier Drew
7950 mutica De Draai Kpbankskloof
7951 mutica Klipbankskloof West
7952 mutica NW Klipbankskloof West
7953 mutica N Klipbankskloof West
7954 mutica Rondeheuwel
7955 mirabilis Van Reenens Crest
7956 mirabilis Van Reenens Crest
7957 mirabilis Van Reenens Crest
7958 mirabilis Van Reenens Crest
7959 mirabilis Van Reenens Crest
7960 mirabilis Postdal
7961 retusa nigra Postdal
7962 floribunda Niekerkshek W STn
7963 floribunda Niekerkshek SW STn
7964 mirabilis W Spitskop De Hoop
7965 mirabilis W Spitskop De Hoop
7966 mirabilis W Spitskop De Hoop
7967 mutica Spitzkop W Rd
7968 mutica Spitzkop W Rd
7969 mutica Spitzkop E Rd
7970 mirabilis Die Kop East(dutoit)
7973a mirabilis NW Die Kop
7973b mirabilis NE Die Kop (Uys =7502)
7974 mirabilis S Klipfontein, Potberg
7975 floribunda Rotterdam W
7976 floribunda N Rotterdam W
7977 mirabilis NW Van Reenens Crest
7978 mirabilis Schuitsberg S
7979 mirabilis Nethercourt
7980 mirabilis Ouplaas, SE Greyton
7982 mirabilis Schuitberg N
7983 marginataXminima S BlueCrane Rest.
7984 mirabilis Droogerivierberg W
7985 mirabilis Droogerivierberg SE
7986 mirabilis Damsekop Montagu
7987 arachnoidea Uitkyk Montagu
7988 mirabilis SE Sandberg Trappieskraalkloof
7989 pumila Lemoenpoort
7990 mirabilis Klipkop Montagu
7991a maculata Top Moddergat E
7991b maculata west above
7992 maculata Hammansberg Radio Stat
7993 maculata Hammansberg top west
7994 maculata Vreesniet, Kanetvlei
7995 herbacea S Brandvlei Brickworks
7996 herbacea E Brandvlei Brickworks
7997 pubescens N Cilmor Cellar
7998 floribunda Kruisriver N
7999 mirabilis Kruisriver N
8000 mirabilis Wegwysersriver
8001 floribunda Kleinkruis, Platkop
8002 maculata/pubescens W Cilmor Cellar
8003 retusa/mirabilis 350m W Bloekombos Rsdal
8004 retusa/mirabilis 50m W Bloekombos
8005 retusa/turgida S Wegwysersriver
8006 floribunda S Wegwysersriver
8007 retusa/turgida 650m W Bloekombos Rsdal
8008 retusa/turgida 1km W Bloekombos
8009 retusa 3km S Riversdale
8010 retusa 8km S Riversdale
8011 pubescens SW point Sandberg
8012 herbacea Klipkranz camp Sandberg
8013 pubescensXherbacea SW point Sandberg
8014 herbacea SE Sandberg, Ribbokkop
8015 herbacea Spes Bona
8016 herbacea N Spes Bona
8017 floribunda Bontebok Park
8018 mutica Sanddrift Drew
8019 maculata Die Nekkies Yt Club
8020 maculata East end Nekkies
8021 maculata 400 W above
8022 maculata 800 W
8023 maculata 1000m W
8024 maculata 1100m W
8025 maculata 1500m W
8026 maculata 2000m W
8027 maculata 500m W resort entrance
8028 mirabilis S Klipport N Bromberg
8029 marginata Bontebok N Pk
8030 mirabilis SW Klipport
8031 mirabilis SWW
8032 pumila Iminga, N Ouhangsberg
8033 herbacea Iminga, N Ouhangsberg
8034 maculata Iminga, N Ouhangsberg
8035 mirabilis Bontebok Pk SW
8036 minima Bontebok Pk SW
8037 minima E of 8035 and 6
8038 venosa Felix Unite
8039 venosa Elsies Camp BNPk.
8040 mirabilis NE Greyton
8041 mirabilis Eshuitsberg E
8042 maculata E Iminga 8034
8043 mirabilis Bontebok N Pk S6644
8044 mirabilis with rossouwii minor
8045 rossouwii ‘minor’ SW t.loc Rooivlei
8046 mirabilis Brakkloof SW
8047 mirabilis Brakkloof SE
8048 mirabilis Rooivlei W
8049 mirabilis Rooivlei N of above
8050 mirabilis Rooivlei NW
8051 mirabilis E. Rooivlei
8052 mirabilis S Welgegund S above
8053 mirabilis S Welgegund S above
8054 arachnoidea Grootkloof, DeDoorns
8055 mirabilis Hammanskraal, Caledon
8056 mirabilis E Uitkyk Greyton
8057 mirabilis Uitkyk, N Breede
8058 mirabilis Wolfkloof Swellendam
8059 floribunda E Wolfkloof Swellendam
8060 mutica Bontebok
8061 marginata Adoonskop rd verge
8062 mirabilis Rietkuil
8063 floribunda W Keurenboom Estate

Cymbiformis Cooperi

I was musing over a whole lot of plants in a nursery all labelled cymbiformis and ignoring the formal classification and recognition of varieties. Gordon Rowley was very critical of my work because he felt that the lesser names were just being dumped. I agree that it is a problem and I was thinking of posting pictures that would facilitate the use of old existing names e.g. planifolia. I was very quickly disenchanted as I looked for pictures.

The field situation regarding cooperi and cymbiformis is as complex as the retusa/turgida/mirabilis/emelyae/pygmaea/mutica issue as well as the fact that each such species assemblage runs over in to still other species. I did google cymbiformis and planifolia to see what the internet had to offer and was amazed how poor and confusing that is. I welcome any questions that cooking up an answer for, may help rationalise and enlighten troubled minds.

The picture is from one population at Kaboega where it is evident to me that there is no distinction between cymbiformis and cooperi, and it gets worse.

There is a problem with the name obtusa as a variant of H. cymbiformis, and I think it, with the name translucence, that really covers the transition between cymbiformis and cooperi. Both those names obtusa and translucens can be applied to a large number of populations that belong in a twilight zone of neither “this nor that”. As I was pondering the other day, plant taxonomy is NOT good science. It is largely treated as a field for nomenclatural rules and recommendations and a stage for intellectual display. There is little connection between the minutae of nomenclature and the reality of what occurs in nature. These pictures are of plants in the Bosberg NE Somerset East and are in a geographical cooperi environment far from the range of cymbiformis. They could fit the concept of “obtusa” as so many other cooperoids do.

Let me add – cymbiformis are those green things that grow as clump formers on steep (rock faces, cooperi are those bluish plants that grow solitarily on the flat. They are the same!!! The names are just at the extremities and the more we know the more names we need??

When I posted those two items about cooper and cymbiformis, I had left gracilis somewhere in the closet. It is just a very difficult pill to swallow that all these amazing variants have (?) to be lumped under one species.

I had another look at the 3 DNA phyllograms in the work by Manning et al, and it can be noted that cooperi, cymbiformis and decipiens all see to pan out in the same lineage. So as I said it is far worse than imagined – although there are some improbables in the various lineages of the phyllograms. But this is just as far as sequencing has got and we have yet to see results from “next generation sequencing”. Here are two images of cooperi from different populations west of Uniondale pass that shocked even myself in saying they are the same species.

Many years ago you gave me a challenge to sort out the cymbiformis cooperi and so off I went to the Eastern Cape; many times. I didn’t understand then how plastic, how variable, the plants were and how rigid the names and descriptions. Some I couldn’t type.

Cymbiformis – Rosette to 130mm φ, partially stemmed, proliferous. Leaves broad ovate to lanceolate, flat to slightly concave, generally <1/3 as thick as wide, usually opaque, green turning yellowish to pink hued on exposure. Inflorescence to 250mm, 10-15 flowers, lax. Flowers white.

Cooperi – Rosette to 120mm φ, often proliferous, stemless. Leaves 20-40, fleshy, swollen, oblong-lanceolate, quickly tapering, acuminate or truncating, marginal spines <2mm long if present. Bluish-green in colour, slightly translucent, with veins usually reddening and leaves developing purplish hues in exposed situations. Inflorescence compact, firm peduncle with many closely arranged flowers, to 20cm long. Flowers 20-30, perianth white.

Maybe it’s time to come back for more field work.

Lawrence Loucka

Yes writing those descriptions was a really funny experience but actually they are remarkably as good as you can get. Cooperi is only distinguishable from cymbiformis by colour, inflorescence and habitat. So what do you do with all those other populations that fall in the small cracks between. The “fieldwork” necessary to establish a workable boundary is to weed out the vast majority of plants that will not fit these artificial strictures of botanical classification and nomenclature. And just what is this stuff we weirdly say is “fieldwork”?

Really! I put it best I think in Update Vol 2 or 3 where I wrote about the need for a starting hypothesis for field work and reporting. I can tell you right now what more unplanned field work will produce – more names and more confusion. I have shown time and again that planned field work shows the interconnectedness of all these “species” we so anxiously claim. But we are also very limited by our perceptions and our expectations. We are taught mechanistically and we are not taught about the illusion of time and space or what reality is or might be. We do not live in a finite fixed world that is the same for each of us. We live within our own perceptions and dreams of what we think is real? We work with definitions of words and things and if we do not agree with each other on these, we will not understand each other. The word “species” is the great catch point hen it comes to biology. Science presently defines it for us by default, as a measure of the similarity of DNA in some sort of sampling (ill-defined and inadequate) process. This is going to create serious problems into the future. The polymath who asked plant taxonomists if they knew what they are doing, is going to have his doubts proven too right – they do not.

Bruce Bayer

I am always on about definition, usually of “species”, but checking up some DNA information reveals that even things such as nucleotides, DNA, and what-not are not actually adequately defined. DNA is said in some meanings, to be the nucleotide chains in the cell nucleus. The nucleotides are said to be the four amino-acids that link the paired strands in the chromosomes. But how are the strands constructed?

Then there are different molecular combinations effectively forming short strands in all tissues and these are also referred to as DNA. The Haworthia sequencing used to establish the recent changes within the aloids is based on up to 7 molecular regions, some of nuclear DNA and some on extra-cellular DNA. It bothers me that the sequencing does not actually tell me very much that I was not aware of simple observing the plants. But I see things in each of the phyllograms for the different regions that persuade me that we have a LOT to learn about DNA sequencing. The lineages for each region are not the same and there are some very weird anomalies that people who do not know the species are clearly not aware of. If they were they would be less confident in their conclusions and their faith in the technology (as it stands to date).

Glad that Lawrence posted that DNA paper Manning et al, because those interested can now see the phyllograms and make up their own minds about what they tell us. My personal experience makes me highly suspicious of all this stuff as even in the paper the authors say they opine that the results give a reliable overview – despite limited sampling and despite the variations in the different lineages the phyllograms illustrate. I would actually like to see phyllograms for each of the 5 DNA regions as well as concatenated ones, for just H. floribunda sampled from many populations. also not just single specimens from each population.

Here we have a case where the plastid sequences do not agree with the nuclear. I also do not like using both regional sequences AND concatenated results. Either concatenation is better or it is not. It seems like the statistical ploy viz. if a test does not give you significance, you keep using different statistical tests until you get one that does. The most critical element is that while these results are used to arrive at a generic classification in the aloids, the exercise was initiated to examine the species relationships in Haworthia only. Had that initial intention been observed (using 3 specimens per population) it could have made a really significant contribution instead of leaving us all in vacuum of opinion and belief.

My present view is that classification has devolved into an intellectual exercise that perpetuates confusion and uncertainty, ensuring endless name changes! Posted (above) is a nice retusaXcooperivenusta‘ hybrid. It does not darken when exposed to direct day-long sunlight.

It is interesting to look at the DNA lineages for H. marxii, H. semiviva, H. pulchella var. globifera. H. mutica and H. marumiana var. archeri. They are, in the three data sets, very conflicting. For me highlight the problem of sequencing where sampling is so limited and based on an identification or classification process itself not based on sequencing. Particularly so when the original intention of the project was to examine that very problem.

This might have been called Haworthia dekenah var. argenteo-maculosa! In my opinion all the retusoids east of the Gouritz river are referable to H. pygmaea. But the truth is that one cannot separate retusa, turgida, mutica, mirabilis, pygmea or even emelyae on the basis of any physically observable evidence.

Indeed there is a problem with correct and consistent naming. The fact is that it can be much like pinning the tail on the donkey. It is all very well having a nomenclatural system that can be a juristic nightmare and an intellectual challenge, but does it work? 

Haworthia cymbiformis is a case in point. According to the system, if a variety is recognised and described, automatically every other variant becomes the variety cymbiformis. This creates an amazing treadmill and we then have to accommodate all those other variants in a practical working system. How does one deal with this when, apart from the problems of one complex system, H. cymbiformis is inextricably tied into H. cooperi.

My conclusion has to be that the system does not work. While there is the fancy intellectual footwork in the upper echelons of botany, there is also the difficulty of dealing with countless variations that are conveniently set aside because they do not fit. Looking at my own attempts to deal rationally with H. cymbiformis and its variants boggles my mind. How did I set the name ‘planifolia’ aside, or renege on an earlier decision to abandon the name ‘obtusa’, and subsequently change my mind again? No matter how one juggles these names, one is faced with the embarrassment of dealing with many more variants that are neither one nor the other of the names one does decide to recognise and use. Let us say that this picture above is typical of H. cymbiformis and then look at some others …

What I tried to do in Haworthia was establish what groups existed that satisfied some rational species definition in also geographic terms. So I abandoned many of the old names for that reason alone. “Planifolia”is a case in point. I could find no clear geographic evidence and recognised the difficulty of actually separating cymbiformis and planifolia as distinct groups. But it is a subjective decision as so many taxonomic decisions are despite the desperate efforts to hide the doubt and the justify them as objective. as you can see from the two pictures, “planifolia’ has broader and flatter leaves rather than the keeled “boathull” shape of cymbiformis. I did not intend that names in synonymy be abandoned and did suggest that such names be considered available for use by collectors and growers.

Quote … ”The location of this note is a powerful reminder that distinctions between species are highly blurred and that alternative solutions are possible”. This answers my question of the previous post and it comes from Haworthia Revisited after tortuous explanations of the varieties in H. cymbiformis. Here is a photo of a cymbiformis in cultivation that could have come from anywhere, including Plutos Vale where H. cooperi var. tenera (?) and H. cymbiformis var. incurvula obfuscate the difference between the two species.

At this late stage in life, I think an honest submission is called for. My life experience with identification of plants is that botanical classification and naming is largely delusional. Taxonomists name things in relation to their experience and seldom get to truly know all the species and variants. Herbarium cupboards hide scores of unusual specimens filed in doubtful places and many unidentified. I have found it very difficult to deal with variations as this posted picture demonstrates. It is a variant of H. cymbiformis that I initially recognised as var. umbraticola in the belief that it substituted for the name var. obesa. At the time I thought that was a name confused with H. cooperi. I did eventually change my mind and recognise obesa as a variant of cymbifomis. But the fact is that cymbiformis is a riverine cliff hanger versus cooperi that is a grassland element.

There fall is a large spectrum of populations occur in different habitats that are neither one nor the other. This situation is also represented in H. retusa and H. turgida where there is further complexity in plant appearance and behavioural character viz. H. mirabilis, H. pygmaea, H. mutica, H. emelyae and more. These are realities of plant classification that taxonomists seem to dismiss or are unable to recognise. Things simply do not follow the tidy branching nature that fancy evolutionary theory, cladograms, phylograms and revisions illustrate.

This is now from an area in the Prince Alfed’s Pass where H. cymbiformis var. transiens, and H. planifolia var. translucens originated. It is to the west of general distribution of where more characteristic H. cymbiformis and H. cooperi occur. But my experience of these two species is that anything that resembles H. cymbiformis is actually a habitat derived variant of H. cooperi, which includes what was a very complex set of variants that I had treated as yet another species viz. H. gracilis. ♦